Nirican
Invertebrates
African Invertebrates 64(3): 221-289 (2023)
DOI: 10.3897/Afrinvertebr.64.111047
Checklist
Checklist of the spiders (Araneae) of South Africa
Ansie S. Dippenaar-Schoeman™, Charles R. Haddad”®, Leon N. Lotz’,
Ruan Booysen®, Rudolph C. Steenkamp*‘, Stefan H. Foord'®
1 SARChI Chair on Biodiversity Value and Change in the Vhembe Biosphere Reserve, Faculty of Science, Engineering and Agriculture, University of Venda, Private
Bag X5050, Thohoyandou 0950, South Africa
2 Department of Zoology and Entomology, University of the Free State, RO. Box 339, Bloemfontein 9300, South Africa
3 Retired from Department of Arachnology, National Museum, P.O. Box 266, Bloemfontein 9300, South Africa
4 The Spider Club of Southern Africa, Bloemfontein, South Africa
Corresponding author: Stefan H. Foord (stefan. foord@univen.ac.za)
OPEN Qaccess
Academic editor: Galina N. Azarkina
Received: 15 August 2023
Accepted: 15 October 2023
Published: 14 November 2023
ZooBank: https://zoobank.org/
EC8097AB-917E-42FB-8B46-
0AA5S71B7636B
Citation: Dippenaar-Schoeman AS,
Haddad CR, Lotz LN, Booysen R,
Steenkamp RC, Foord SH (2023)
Checklist of the spiders (Araneae) of
South Africa. African Invertebrates
64(3): 221-289. https://doi.
org/10.3897/Afrinvertebr.64.111047
Copyright: © Ansie S. Dippenaar-Schoeman et al.
This is an open access article distributed under
terms of the Creative Commons Attribution
License (Attribution 4.0 International -
CC BY 4.0).
Abstract
Achecklist of 2265 spider species and subspecies, 495 generaand 71 families is provided.
Data were extracted from the South African National Survey of Arachnida database and
over 200 taxonomic revisions, ending December 2022. Global distributions, endemicity
and conservation assessment using IUCN Criteria are provided for each species. A total
of 1325 spp. are endemic to South Africa (58.5%), 126 spp. (5.6%) are of special concern
and 693 spp. (30.6%) are Data Deficient (DD), while 15 species were described without
exact locality data. Most species (1444 spp., 63.8%) are widely distributed with no known
threats and are of Least Concern. A total of 1316 spp. (57.6%) are known from both
sexes and 23 spp. (1.0%) were described from juveniles. Salticidae is the most species-
rich family (354 spp.), followed by Gnaphosidae (195 spp.), Thomisidae (143 spp.) and
Araneidae (100 spp.) and ten families are represented by a single species.
Key words: Conservation status, endemicity, faunistic surveys, global distribution, pro-
vincial, sample bias, SANSA, South African National Survey of Arachnida
Introduction
The emerging field of conservation biogeography concerns species’ distribu-
tion dynamics and how they relate to biodiversity conservation (Robertson et
al. 2010); and its main currency is valid species-level determinations and dis-
tribution data. However, the knowledge and insights gained from these data
depend on whether species are valid and systematically established, the exis-
tence of clear diagnoses to aid positive determinations, documented biology
and reconstructed evolutionary histories. These activities depend on the prima-
ry data collected with specimens that provide vast information (Hamer 2012).
Therefore, curating and disseminating this information in retrievable formats,
such as a national list, is essential.
The South African National Survey of Arachnida (SANSA), which was initiat-
ed in 1997 (Dippenaar-Schoeman et al. 2015), had the main aim of document-
ing the arachnid fauna of South Africa by collating all the data into a relation-
al database. The second phase of SANSA (2006-2010), funded by the Royal
221
Ansie S. Dippenaar-Schoeman et al.: South African spiders
Norwegian Ministry through the Threatened Species Programme of the South
African National Biodiversity Institute (SANBI), was critical, as it surveyed large
under-represented parts of South Africa (Dippenaar-Schoeman et al. 2015), cul-
minating in the release of an unpublished report: the First Atlas of the Spiders
of South Africa (FASSA) in 2010 (Dippenaar-Schoeman et al. 2010). Until now,
this Atlas was the only known national list of South African spiders.
Only afew African countries have published spider species checklists: Botswana
(Eagle 1985), Namibia (Griffin and Dippenaar-Schoeman 1991), Zimbabwe
(FitzPatrick 2001), Sudan (Dunlop and Siyam 2014), Tanzania (Russell-Smith
2020) and Kenya (Kioko et al. 2021). Here, we provide the first annotated spider
checklist for South Africa, based on published data and accessioned specimens
in collections up to the end of December 2022. We also include references to
unpublished resources that add value to the data and results summarised here.
Material and methods
Study area
The area covered is South Africa (Fig. 1A), excluding the enclave of Lesotho
and Eswatini. It includes the following nine Provinces: Eastern Cape (EC), Free
State (FS), Gauteng (G), KwaZulu-Natal (KZN), Limpopo (L), Mpumalanga (M),
Northern Cape (NC), North West (NW) and Western Cape (WC).
Source of information
As part of SANSA, all available data were collated in a relational database main-
tained by the Agricultural Research Council (Dippenaar-Schoeman et al. 2012).
This database is not publicly accessible, but the data retrieved from this database
and used for this manuscript are available online in a tidy format (https://zenodo.
org/record/8249595). These data were used to identify the gaps in the geograph-
ical coverage and guide the field surveys of phase II of SANSA, which were con-
ducted using a standardised protocol (Haddad and Dippenaar-Schoeman 2015).
All the specimens from postgraduate ecological surveys from 12 universities in
South Africa were deposited in the National Collection of Arachnida (NCA) at the
Agricultural Research Council in Pretoria as voucher specimens. Many specimens
were also received from the public participating in SANSA surveys, including
members of the Spider Club of Southern Africa and Reserve managers. A large
number of specimens were collected during national and international projects
included in unpublished studies, but included in SANSA newsletters. However,
these newsletters went through an editorial process and most are available online
(Suppl. material 1). Voucher specimens of sampled material were also deposited
National Museum in Bloemfontein, Ditsong National Museum of Natural History
in Pretoria and KwaZulu-Natal Museum in Pietermaritzburg.
Checklist structure
The checklist is based on data extracted from the SANSA database and ref-
erences to the taxonomic and more than 200 published/unpublished faunis-
tic surveys up to the end of December 2022 (Suppl. material 1). Only spiders
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 229
Ansie S. Dippenaar-Schoeman et al.: South African spiders
identified to species level are included. Families and species are arranged al-
phabetically. For each species, we provide:
Endemicity
The following terms are used: SAE: species endemic to South Africa;
STHE: species endemic to southern Africa; AE: species endemic to the
Afrotropical Region; C: species that also occur beyond the Afrotropical Region.
Distribution
The following terms are used: 6: species is known only from the type locality in
South Africa; 5: species is known from several localities in the same province as
the type locality; 4: species were sampled from two adjacent provinces; 3: species
were sampled from 2 three provinces in South Africa; 2: species occurs outside
South Africa, but within southern Africa; 1: Afrotropical Region; 0: cosmopolitan.
Provincial data
Species listed per South African province obtained from primary label data of
collected specimens, as reported in taxonomic and faunistic publications and
the SANSA database.
Conservation assessment
As part of the Spider Red Listing Project (Foord et al. 2020), the preliminary con-
servation status of species as determined are listed with the following codes:
DD (Data Deficient): species usually known from only one sex or based on old
material without detailed locality data and where the species is difficult to iden-
tify (DDT: Data Deficient due to Taxonomic reasons); LC (Least Concern): spe-
cies with a broad distribution (categories 0-2), without known threats; those
of categories 3 and 4 are South African endemics (SAE) and many of them are
also LC. Species of special concern (Rare, Critically Rare, Vulnerable, Threat-
ened) usually belong to categories 5 or 6.
History
Almost a third of South African species were described from 1900-1920
(739 spp.) (Fig. 1B), focusing largely on the fauna of the coastal provinces, as
most practising arachnologists were stationed there. From 1960 to 1980, there
was a considerable decline in the description of new species due to lack of
practising taxonomists. However, during the last three decades there has been
a resurgence; more than 700 species have been described since 1980, mainly
due to new appointments, modern taxonomic revisions, the training of several
South African taxonomists and the efforts of taxonomists overseas. Ongoing
SANSA surveys continued contributing accessions at the NCA and specimen
identifications have also contributed to a constant increase of new species re-
cords (Fig. 1B). The increase in taxonomic output since 1997 can partly be
attributed to SANSA (Dippenaar-Schoeman et al. 2015).
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 293
Ansie S. Dippenaar-Schoeman et al.: South African spiders
South African species
1700-1799 ;
1921-1940 -—— —
1941-1960
1961-1980
1981-2000
2001-2022
Figure 1. A Map of South Africa and its nine provinces B temporal trends in species described from South Africa (1700-2023).
Species identification
Unfortunately, the original descriptions of many species described from 1700-
1950 are often rudimentary, only from one sex, lacking detailed morphological
data and without drawings or detailed collecting data. Most type specimens
are also housed in overseas museums or are lost. One of the most significant
constraints for SANSA is the lack of taxonomic revisions for many of the larger
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 224
Ansie S. Dippenaar-Schoeman et al.: South African spiders
spider families in South Africa, with reference to Agelenidae, Araneidae, Dic-
tynidae and Theridiidae. The number of South African species will inevitably
increase, aS many specimens housed in collections remain unidentified due to
lack of taxonomic resolution.
With all the available data, SANSA has developed online photo identi-
fication guides for the 72 families to address the taxonomic constraints
(Suppl. material 2). These guides list known information for all genera and spe-
cies listed from South Africa. Species-level information includes a distribution
map for South Africa, drawings (if available) and photographs of diagnostic
characteristics, notes on their behaviour, a conservation assessment and pos-
sible threats. Complete guides can be downloaded from the World Spider Cat-
alogue, as well as from Zenodo (https://zenodo.org/communities/sansa/) (for
complete list see Suppl. material 2).
Results and discussion
A total of 2265 spider species and subspecies, belonging to 495 genera and
71 families, are presently known from South Africa (Table 1). The Salticidae
(354 spp.), Gnaphosidae (195 spp.), Thomisidae (143 spp.) and Araneidae (100
spp.) are the most species-rich families. A total of nine families are only repre-
sented by a single species.
Table 1. Spider families from South Africa listing the number of genera and species
per family, as well as the number endemic to South Africa (SAE), the number of Data
Deficient (DD) species, the species of Least Concern (LC) and the number of species of
Special Concern (SC).
FAMILIES GEN SPP SAE DD LC Sc
Agelenidae 5 10 0 0 10 0
Amaurobiidae 5 16 16 11 3 2
Anapidae 3 4 3 2 1 1
Anyphaenidae 1 1 0 0 1 0
Araneidae 40 100 20 3 90 3
Archaeidae 1 14 14 6 2 6
Atypidae 1 2 2 0 0 2:
Barychelidae 2 3 0 0 3 0
Bemmeridae 2 33 32 30 3 0
Caponiidae 3 16 10 6 8 2
Cheiracanthiidae 3 47 36 20 26 1
Cithaeronidae 1 1 1 1 0 0
Clubionidae 1 26 le, 12 14 0
Corinnidae 15 44 22 11 28 5
Ctenidae 2 8 8 1 Zz 0
Cyatholipidae 6 16 16 4 7 5
Cyrtaucheniidae 1 29 28 17 12 0
Deinopidae 2 7 1 0 6 0
Desidae 2 2 0 0 2 0
Dictynidae 4 5 2 1 4 0
Drymusidae 1 5 5 1 0 4
Dysderidae 1 1 0 0 1 0
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 295
Ansie S. Dippenaar-Schoeman et al.: South African spiders
FAMILIES
Entypesidae
Eresidae
Euagridae
Filistatidae
Gallieniellidae
Gnaphosidae
Hahniidae
Hersiliidae
Idiopidae
Ischnothelidae
Linyphiidae
Liocranidae
Lycosidae
Microstigmatidae
Migidae
Mimetidae
Miturgidae
Mysmenidae
Nephilidae
Nesticidae
Oecobiidae
Oonopidae
Orsolobidae
Oxyopidae
Palpimanidae
Philodromidae
Pholcidae
Phyxelididae
Pisauridae
Prodidomidae
Pycnothelidae
Salticidae
Scytodidae
Segestriidae
Selenopidae
Sicariidae
Sparassidae
Stasimopidae
Symphytognathidae
Telemidae
Tetragnathidae
Theraphosidae
Theridiidae
Thomisidae
Trachelidae
Trochanteriidae
Uloboridae
Zodariidae
Zoropsidae
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
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Provincial surveys
The number of families, genera and species for each of the nine provinces
are listed in Table 2. KwaZulu-Natal has the highest number of known species
(1024), followed by the Western Cape (908) and Limpopo Province (905), while
North West had the least (344). Note: almost all the surveys without citations
listed below, can be found in the grey literature, references and DOls to these
documents being included in Suppl. material 1 of the supplementary material.
Eastern Cape (Fig. 2A): The Eastern Cape becomes progressively wetter
from west to east. The west is mostly semi-arid Karoo, except in the far south,
where there is a temperate rainforest in the band between the Cape Fold Moun-
tains and the Indian Ocean. The coast is generally rugged with interspersed
beaches. Most of the Province is hilly to very mountainous to the west of East
London and Queenstown and towards the KwaZulu-Natal border - a region
known previously as Transkei — it is a lush grassland on rolling hills, punctuated
by deep gorges with intermittent forest. A total of 837 species (137 endemic)
have been recorded from the Province (Table 2).
Major surveys in this Province include: Addo Elephant National Park (Dippe-
naar-Schoeman et al. 2020); Amathole Mountains (Haddad et al. 2023); Asante
Sana Private Game Reserve; Jeffreys Bay; Mkambati Nature Reserve (Dippe-
naar-Schoeman et al. 2011); Mountain Zebra National Park (Dippenaar-Schoeman
1988, 2006); Nduli and Luchaba Nature Reserves (Niba and Yekwayo 2016); Sila-
ka Nature Reserve (Forbanka and Niba 2013); Thyspunt; Tsolwana Game Reserve.
Free State (Fig. 2B): The Free State is situated on the central escarpment and
is comprised of a succession of flat grassy plains sprinkled with pastures and
cultivated lands, resting on a general elevation of 1200 m, only broken by the
occasional hill. The rich soil and pleasant climate allow for a thriving agricul-
tural industry. It is known locally as the breadbasket of South Africa with more
than 30,000 farms, which produce over 70% of the country’s grain.
The Free State is the third largest province (10.6%) and a total of 517 species
(25 endemic) have been recorded here (Table 2).
Major surveys in the Province include: Amanzi Private Game Reserve (Haddad
and Butler 2018); Farm Bankfontein, Luckhof District (Haddad 2021); Farm Deel-
hoek, Bloemfontein District (Haddad and Dippenaar-Schoeman 2002, 2006a;
Haddad 2005); Farm Lusthof, Edenville District (Dippenaar-Schoeman et al. 1978);
Table 2. Number of families, genera, species and endemics recorded for each of the nine provinces in South Africa, as
well as the percentage of all the known South African species.
Provinces
Eastern Cape
Free State
Gauteng
KwaZulu-Natal
Limpopo
Mpumalanga
North West
Northern Cape
Western Cape
Family
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60
57
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Genera Species Endemics % spp. total SA
312 837 137 37
224 517 25 24
250 563 11 25
362 1024 159 45
346 905 55 Al
279 645 20 28
188 344 6 15
236 559 62 25
319 908 261 41
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 297
Ansie S. Dippenaar-Schoeman et al.: South African spiders
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Figure 2. Spatial distribution and number of accessions for localities in: A Eastern Cape B Free State C Gauteng
D KwaZulu-Natal E Limpopo.
Erfenisdam Nature Reserve (Fourie et al. 2013; Haddad et al. 2015); Florisbad Re-
search Station (Lotz et al. 1991); Free State National Botanical Gardens (Butler
and Haddad 2011; Neethling and Haddad 2013, 2019; Haddad et al. 2019, 2021;
Luwes and Haddad 2020); Golden Gate Highlands National Park (van der Merwe
et al. 2020); Kalkfontein Dam Nature Reserve, and Mpetsane Conservancy.
Gauteng (Fig. 2C): This is the smallest province, covering only 1.4% of South
Africa (Fig. 2C) and falls within both the Savannah and highly threatened Grass-
land Biome with approximately 83% of the Province falling within the Highveld
Grassland vegetation type, of which a mere 0.8% is currently conserved in
South Africa. A total of 563 spp. (11 endemic) have been recorded from this
Province (Table 2).
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 228
Ansie S. Dippenaar-Schoeman et al.: South African spiders
Major surveys in this Province include: Bakwena Cave in Irene (Durand et
al. 2012); Ezemvelo Nature Reserve; Faerie Glen Nature Reserve in Pretoria;
Gauteng caves (Dippenaar-Schoeman and Myburgh 2009); Groenkloof Nature
Reserve; Irene survey; Kliprivierberg Nature Reserve; Rietondale Research Station
(Van den Berg and Dippenaar-Schoeman 1991); Roodeplaat Research Station
(Dippenaar-Schoeman 1976; Roodeplaatdam Nature Reserve (Dippenaar-
Schoeman et al. 1989; Engelbrecht 2013); Pretoria National Botanical Gardens;
Serene Valley; Suikerbosrand Nature Reserve; Tswaing Crater Nature Reserve;
and Waterkloof Nature Reserve.
KwaZulu-Natal (Fig. 2D): KwaZulu-Natal covers 7.6% of South Africa and har-
bours many vegetation types. Grassland predominates in the western interior
of the Province, with savannah towards the east. Alpine vegetation occurs on
the high Drakensberg Escarpment on the border with Lesotho. The Coastal Belt
vegetation is regularly interrupted by large river systems, where a thicket-type
vegetation is often present in the valleys. A total of 1034 species have been
recorded, which constitutes 45.3% of the spiders of South Africa (Table 2).
Dr. Reginald Lawrence, one of the best-known South African arachnologists,
was Stationed at the Natal Museum from 1935 to 1986 and he extensively sam-
pled and described species from the Province.
Major surveys in the Province include: Buffelsdraai Landfill Conservancy;
Howick, Farm Wakefield; Drakensberg; False Bay Park (Lovell et al. 2007);
Hluhluwe Game Reserve (Mgobozi et al. 2008); Farm Vergeval, Pongola district
(Dippenaar-Schoeman et al. 1978); iSimangaliso Wetland Park, various localities;
Ithala Game Reserve; Kosi Bay Nature Reserve; Maputaland (Lawrence et al.
1980); Maluti-Drakensberg Transfrontier Project (MDTP); Midlands, Good Hope
and Maybole plantation estates (Yekwayo et al. 2016, 2017); Midlands, Ncandu
Falls, Moorfield Mountain Farm and Normandien Farms (Booysen and Haddad
2021); Ngome State Forest (Van der Merwe et al. 1996; Dippenaar-Schoeman
et al. 2006); Ndumo Game Reserve (Haddad et al. 2006; Haddad 2016, 2022);
Ophathe Game Reserve (Haddad and Dippenaar-Schoeman 2015); Vernon
Crookes Nature Reserve; Phinda Game Reserve (Lovell et al. 2010); Richards Bay
rehabilitated coastal forest (Dippenaar-Schoeman and Wassenaar 2002, 2006);
Sani Pass altitudinal gradient; Sodwana Bay National Park; Tembe Elephant Park
(Haddad et al. 2010); and uMkhuze Game Reserve (Lovell et al. 2007).
Limpopo (Fig. 2E): This Province covers 10.6% of South Africa. The vegeta-
tion ranges from Afromontane forests, savannahs, shrubland, to semi-desert
areas with small trees and bushes. The landscape also ranges from mountain-
ous to flat plains. The first surveys in the Province were undertaken during a
five-year harvester termite control project (Dippenaar-Schoeman et al. 1978).
Another large dataset became available during surveys looking at the effect of
chemical control of quelea finches on the Springbok Flats.
A total of 905 species (55 endemic) is known from the Province (Table 2).
Major surveys in the Province include: Atherstone Game Reserve; Blouberg
Nature Reserve (Muelelwa et al. 2010; Foord et al. 2019); Farm Amsterdam,
Dendron District (Dippenaar-Schoeman et al. 1978); Farm Zandrivier, Lephalale
District, Bioblitz; Ka-Ndengeza Village (Joseph et al. 2017; Foord et al. 2018);
Klein Kariba Resort; Lekgalameetse Nature Reserve (Foord et al. 2016); Leph-
alale; Limpopo Valley; Luvhondo Nature Reserve (Foord et al. 2008); Makelali
Nature Reserve (Whitmore et al. 2001, 2002); Marakele National Park; Nylsvley
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 229
Ansie S. Dippenaar-Schoeman et al.: South African spiders
Nature Reserve (Heidger 1988; Dippenaar-Schoeman et al. 2009); Pietersburg
Nature Reserve (Dippenaar et al. 2008); Rust de Winter cotton surveys (Dip-
penaar-Schoeman et al. 1999b); Soutpansberg long-term survey (Foord et al.
2002); Sovenga Hill (Modiba et al. 2005); Syferkuil research site; Venetia Lim-
popo Nature Reserve; Vhembe Biosphere Reserve; and Waterberg Biosphere.
Mpumalanga (Fig. 3A): Mpumalanga covers a total of 6.5% of South Africa's
surface. The province contains several distinct physiographic regions: the High-
veld, a plateau ranging in elevation from 1200 to 1800 metres in the west; the
forested Drakensberg Mountains rising to more than 2300 metres in the east;
and the Lowveld, a bush-clad plain that slopes gently upward towards the Leb-
ombo Mountains on the Mozambique and Eswatini borders to the east. Much
of Mpumalanga is drained by eastward-flowing tributaries of the Limpopo River.
A total of 645 species (20 endemic) have been recorded from the Province
(Table 2).
Major surveys in the Province include: Barberton Nature Reserve (Mwabvu
and Yekwayo 2019; Mavasa et al. 2023); Delmas (Bt maize); Kruger National
Park (Dippenaar-Schoeman and Leroy 2003; Jonsson et al. 2010; Robertson et
al. 2011), Lowveld Botanical Gardens, Mariepskop (Taylor et al. 2020); Marble
Hall (Bt cotton) (Mellet et al. 2006); Mbombela (avocado, macadamia, citrus)
24.0°S 5
24.5°S J 26°S 5
25.0°S 4
Accession: Accessions
500 28°84
25.5°3 4 400 200
300 150
26.0°S4 200 100
100 50
30°S 4
26.5°S4
27.0°S 7
32°S 4
27.5°S 5
T T T T T T T T
28.5°E 29.0°E 29.5°F 300°E 30.5°E 31.0°E 31.5°E 32.0°E 16°E 18°E 30°F 22°F DAE 26°F
24.5°S 4
31°S
25.0°S 4
Accessions
25.5°3 | Accession 33-5
300
26.0°5 4 300
200
26.5°S 5 200 330g
100
100
27.0°S 4
27.5°3 5 34°S
28.0°S 4
23°E 24°E 25°E 26°E are Ore. 35°S
Figure 3. Spatial distribution and number of accessions for localities in: A Mpumalanga B Northern Cape C North West
D Western Cape.
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 230
Ansie S. Dippenaar-Schoeman et al.: South African spiders
(Van den Berg et al. 1992; Dippenaar-Schoeman 1998; Dippenaar-Schoeman et
al. 2001a, b; Dippenaar-Schoeman et al. 2005a); Steenkampsberg (Jansen et
al. 2013); and Sterkspruit Nature Reserve (Mwabvu and Yekwayo 2019).
North West (Fig. 3C): This Province covers 9.5% of the surface of South
Africa. Much of the Province consists of flat areas of scattered trees and
grassland. The Magaliesberg mountain range in the north-east extends about
130 km from Pretoria to Rustenburg. The Vaal River flows along the southern
border of the Province.
A total of 344 species (six endemic) are known from this Province (Table 2).
The only long-term survey (4 years) was undertaken in the Kgaswane Nature
Reserve. Students at the North-West University and University of Johannesburg
participated in agro-ecosystem surveys focusing on cotton near Brits and Bt
maize near Potchefstroom (Botha et al. 2015). Other surveys were mainly
funded by SANSA and the Spider Club: Barberspan Nature Reserve; Borakalala
Nature Reserve; Pilanesberg National Park; and Zeerust.
Northern Cape (Fig. 3B): The Northern Cape is South Africa’s largest Province
(29.7%). The Karoo Basin dominates this Province and consists mainly of sedi-
mentary rocks and some dolerite intrusions. The south and south-east of the Prov-
ince are high-lying, 1200-1900 metres, in the Roggeveld and Nuweveld Districts.
The west coast is dominated by the Namaqualand region, famous for its spring
flowers and Succulent Karoo vegetation. This area is hilly to mountainous and con-
sists of granites and metamorphic rocks. The central areas are generally flat with
interspersed salt pans. Kimberlite intrusions punctuate the Karoo rocks, giving the
Province its most precious natural resource, diamonds. The north is primarily Kala-
hari Desert, characterised by parallel red sand dunes and acacia tree dry savannah.
In spite of being the largest province, only 559 species (62 endemic) have so
far been recorded in this Province (Table 2).
Very few sites have been intensively sampled: Akkerendam Nature Reserve;
Augrabies Falls National Park; Brand-se-Baai; Diamond Route Reserves
(Benfontein, Dronfield, Rooipoort and Tswalu Game Reserves; Koingnaas;
Namaqua National Park; Namaqua strip mining; Prieska pistachio orchards
(Haddad et al. 2004, 2005; Haddad and Dippenaar-Schoeman 2005, 2006b;
Haddad et al. 2009); Nigramoep; Richterveld National Park; Tankwa Karoo
National Park; and Witsand Nature Reserve.
Western Cape (Fig. 3D): This Province covers 9.7% of the surface of South
Africa. Spiders were included in the Cape Nature Western Cape State of Biodi-
versity 2012 report (Veldtman 2012), a first for South Africa.
A total of 938 spp. are known from the Province representing 41.1% of South
African species. Major surveys include: Aardvark Nature Reserve; Beaufort West;
Borrelfontein; Bontebok National Park; Cape Peninsula (Sharratt et al. 2000);
Cape Floristic Region (Pryke and Samways 2008, 2009, 2010, 2012; Theron et al.
2020a, 2020b); Cape Winelands (Gaigher and Samways 2010, 2014; Geldenhuys
et al. 2021, 2022); Coast to Karoo transect: Spiders of the Cederberg (Foord and
Dippenaar-Schoeman 2016); De Hoop Nature Reserve (Haddad and Dippenaar-
Schoeman 2009); Fernkloof Nature Reserve; Jonkershoek (Swart et al. 2017);
Karoo National Park (Dippenaar-Schoeman et al. 1999a); Kirstenbosch National
Botanical Garden (Tucker 1920); Kogelberg Biosphere Reserve between
Kogelberg and Protea survey (Coetzee et al. 1990; Visser et al. 1999; Yekwayo et
al. 2018); Robben Island (Mukherjee et al. 2010; Roets and Pryke 2013; Cooper
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 231
Ansie S. Dippenaar-Schoeman et al.: South African spiders
et al. 2017); Southern Cape Forest Complex (Swart et al. 2018, 2020); Swartberg
National Park (Dippenaar-Schoeman et al. 2005b); Table Mountain National
Park; Tierberg Long-Term Ecological Research site (Dean 1988; Dean and Milton
1995; Henschel and Lubin 2018); and Tierhoek Organic Farm (Arvidsson et al.
2020).
Conservation
Many species recorded from South Africa (946 spp., 41.4%) are still only known
from one sex. Only 1316 spp. (57.6%) are known from both sexes and 23 spp.
were described from juveniles (Table 4), resulting in many species that are still
Data Deficient.
Conservation status
The majority (1445 spp., 64%) of species have a wide distribution without
known threats and are listed as Least Concern (Table 3). Sufficient data is still
lacking for 691 spp. (31%), which could not be evaluated and are listed as Data
Deficient; 504 of these species are data deficient for taxonomic reasons, 187
are lacking sufficient distribution data to make informed assessments and 288
(12.6%) are known only from the type locality. Only 129 spp. (5.6%) are of spe-
cial conservation concern (Table 3).
Table 3. Conservation status and endemicity of the spider species sampled from South
Africa.
CONSERVATION STATUS NO SPP %
Data Deficient (DD) | 692 31
Not Evaluated (NE) 2 0.001
Least Concern (LC) 1445 64
Special concern 126 5.6
* Rare 43 2
* Critical rare (CR) 22 1
* Endangered (EN) 24 1
* Vulnerable (VU) 32 1.4
* Near threatened (NT) 5 0.2
Endemicity
0 — Africa and wider (C) 86 | 3.8
1 — African endemics (AE) 460 20.4
2 — Southern African endemics (STHE) 394 | 17.4
3-5 South African endemics (SAE) 932 41.3
6- Known only from type locality 390 17.3
Endemicity
Almost the two-thirds of recorded species (1322 spp., 58.4%) are endemic to
South Africa (Table 3; Suppl. material 2), while 3.8% have a distribution beyond
the Afrotropical Region, including several cosmopolitan species, 460 spp.
(20.4%) which are endemic to Africa (AE) and 394 spp. (16.9%) only known
from southern Africa.
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 232
Ansie S. Dippenaar-Schoeman et al.: South African spiders
Table 4. Checklist of spiders (Araneae) of South Africa.
‘ [Gow stars evoewicr|PROV|EC|FS|6 KAN] LM) NC NW] We
FAMILY AGELENIDAE C.L. Koch, 1837
‘Ageenaassralesimen 96 «tt | | da] | a fa]
Agen goedesiRowwert955 2) SET ta
Perotiedesetcole(Smon 191) SE
2 THE
Olorunia punctata Lehtinen, 1967 LC AE
Tegenaria domestica (Clerck, 1757)
a
F
B
B
F
B | Mistaria zuluana (Roewer, 1955)
M
B
B | Tegenaria pagana C.L. Koch, 1840
B
| Se eh | | = |
Ae |
=
a
Tegenaria parietina (Fourcroy, 1785) pelt iis i
Chresiona convexa Simon, 1903
Chresiona invalida (Simon, 1898)
Chresiona nigrosignata Simon, 1903
Chumma bicolor Jocqué & Alderweireldt, 2017
Chumma foliata Jocqué & Alderweireldt, 2017
Chumma gastroperforata Jocqué, 2001
Chumma inquieta Jocqué, 2001
Chumma interfluvialis Jocqué & Alderweireldt, 2017
‘hrs ster ocque&Aldeweels, 2017 OSA
‘hrm shone oceueeAllerveke, 20176 svoT Se ET
Macobinunceter(Smon 1099) SOOT SAE we
Obst armeteLeninen 1967 «S| OTSA WE
Pseudowinusanouaus Puce 1908 OTSA
Pseudowinuspolias Puce 1904S
ie
il
F
F
F
F
B
B
B
B
B | Chumma Striata Jocqué & Alderweireldt, 2017
B
M
F
F
M
B
F
Pseudauximus reticulatus Simon, 1902 | pot | sae | wee | |
[Crozetulus hodesiensis Brgnol 1981 [2 | tC |THE |||
Crozetulus scutatus (Lawrence, 1964) [8] CR | SAE | wee | | | |
=
Acusiias africanus simon,1895 tt] tt tt |
JAethriscus olivaceus Pocock 1902 |] te] ET ttt tft) |
[Afracantha camerunensis (Thorell 1899) || tc || ty] | ft tia} |
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 233
Ansie S. Dippenaar-Schoeman et al.: South African spiders
' [conSTaTUS| eNOEMIITY PROV EC|FS|6/ KAN] LM) NC [NW | We
F[AgepeawecncaPocok te SCS~;‘ ‘(CSA
€ Agopeausrls Wolckene, 08) A
€ Argopeavpabis(ucs 186)
g AgopeliBiomt97 it wT
g Agopertoe abst a
€ Agopewtociaatfossil7)
€ Agopetphokinodon 1997
€ Bjooraneis legen (resshottecimunds 1979) AE
F msvisconicraPocok T9022) SMES Ta
6 cooesvissecvpioeFabiows 155) A
€ Concsvisthoneze Gregan 6 RAE
€ cowwstisvone Bechval 666 wt
€ Oadometsatemaninewi1923 SNE OMNES
F OadometeedeberRo&OppenaarSchoomen 2008 |S) EN SAE OMNE T
F Oadometo tropes OPambndee te) 1)
G ctaerorereckunne.206 tw
€ eoswiaoweoere)
€ cplos outa Walerser 160
€ Ophaerousinaus(smonst) AE
€ rancmenades mente) wt
8 Orophorowcan Fossil) a
F yrephorpaosikaschie7 2) ME |
g eronocrcaba(Smon tess) Ow
F cestcantefabeonbeute i679
€ estrocanemivoestuterte73 tw
€ estrcanesorgureenac toch eA
€ estracante verso Walkeneer 1682) A
F estonabewienert 73a) EL
G ceoimcootugen 10
€ Mpsoconecrcmacuataahlt) AE
€ Hypsoange ovaries 199) 2) STE
© HpsoangelyplanoiesCoporacca 947 AE
e Hpsooneenyomaca Suntan)
F mecaratnsversmoni9o SAE
B eccratiqutesmonions AE
e woye covense(CLKoehTe@) A
B
B
F
B
F
B
F
B
B
B
B
B
M
F
Isoxya mossamedensis Benoit, 1962 ea | 1
Isoxya mucronata Walckenaer, 1841
Isoxya stuhImanni Bosenberg & Lenz, 1895
Isoxya tabulata (Thorell, 1859)
lsoxya yatesi Emerit, 1973
Kilima decens (Blackwall, 1866)
Larinia bifida Tullgren, 1910
o
Larinia chloris (Audouin, 1826)
Larinia natalensis (Grasshoff, 1971)
— | — | —
Lipocrea longissima (Simon, 1881)
Mahembea hewitti (Lessert, 1930)
Megaraneus gabonensis (Lucas, 1858)
Nemoscolus cotti Lessert, 1933
-
i?)
°
Nemoscolus elongatus Lawrence, 1947
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 234
Ansie S. Dippenaar-Schoeman et al.: South African spiders
' [consTaTUS| ENDEMIC PROV |EC|FS|6|KAN| |W) NC [NW | We
NanoscousebsomeSmon 1897 ———S~*~SS (SCTE
Nanoscouswbcoe(Smon tee) 2) tHE TT Tt
Nanos ghtbuncausSimon 872] tHE
Nenespzacorspeitasinon 19038) OTE
Nesscomabenisierd) tf
Nesscomonguowa(scbeweli97)
Newscom bondeiSimen tee) tt Tt
NesscoachornPave. 086)
Newscomnina(CLkochie) tf
Nesscoaknersis Grossing)
Newscom mort Veo 188) 8) ta
Nessconnoveleiment90)
Nesscoapencpes(aschT6) 1) wt
Neoscom quaarobtose Gossho1986 ot]
NesscomqurcooRobens.198 St)
NessconaropieTieh 169) A
Necscomrfpabistuces 198) tf
Nessconasubisea(CLKochI@) Ot
B
B
B
B
B
B
B
j
B
B
B
B
B
B
B
€ Neoscontrengue Kosei 188)
€ Neosconviions ecw 188)
€ Nephingsorenatefobicus75)
M
B
B
B
B
B
B
B
B
B
B
B
—
— | —_
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—
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BE
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—
—_—
Paplon dorebiessents55) SSS ttt
Papletorethronilckval 65) Tt
Pacpleonewoler(Boowal 188) tHE
Pacronesonescopis Poco 1686) Tt TTT
Parroessperoraus(Thoeh 689)
ParroessspecarrtKrent665)
Paslotus dppeaneRotteHoddod 25S OTEK
Patystuclersmon 198 tA
Pasoncoabolmbaasinon ®t
Prsonicanigataenaa(Simen 1909) tt
Pasoncaceroesmon 185 tA
Prensconatrouus(Fobieus 76) tHE ttt
Shgsabodomworout950 St
Shgobwenceitecowt990) tt
‘ShgatoypamandloKurnerSHomiga, 20028 tA
E
FAMILY ARCHAEIDAE C. L. Koch & Berendt, 1854
B
B
B
F
M
B
F
—_—
—_
—
ee
foal
ci
if
fia
=
vi
Fil
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 235
Ansie S. Dippenaar-Schoeman et al.: South African spiders
F | Afrarchaea harveyi Lotz, 2003
F | Afrarchaea kranskopensis Lotz, 1996
M_| Afrarchaea lawrencei Lotz, 1996
B | Afrarchaea neethlingi Lotz, 2017
B | Afrarchaea ngomensis Lotz, 1996
F | Afrarchaea royalensis Lotz, 2006
B | Afrarchaea woodae Lotz, 2006
FAMILY ATYPIDAE Thorell, 1870
M | Calommata meridionalis Fourie, Haddad & Jocqué, 2011
oO
B | Calommata transvaalica Hewitt, 1916
FAMILY BARYCHELIDAE Simon, 1889
F | Pisenor arcturus (Tucker, 1917)
B | Pisenor notius Simon, 1889
-
_
i?)
F | Sipalolasma humicola (Benoit, 1965
FAMILY BEMMERIDAE Simon, 1903
F [Homostol aberetyi(Pucel,t905) —==SSS« S| UOT «| «SAE | ROE aS
F Hemost pana Henit1919) 8S i
F | Homostola reticulata (Purcell, 1902) fs| pot | sae |wee| | | Lie ae il
F (Homostole ibecuaSimon 1928) tS
B HomostezebinaPucel, 1902S
M SproctenusermeusHewit.ii9SSSS=S| SCOTS ET
F SprecenusbroomiTusker 1917) OTSA WOE
B Sprectenscambiewe Puce 902) 6 ODS WE
M Sprecenuscoenieuslawence 19526) OTSEGO
8 Sprectenscoinus(Pocock 1900S OSE WE
€ SproctenwscuvpesHewit,1919—SSSSSS«|SODSSSCSESCKNE ST
M SprectenseniisLawence,1958 8) SAE KING
F Sprectenus avopunctows Puce. 1903) 6) OT SAE ET
F Sprocenusfossarus(Poook 1900) 6S ET
F Sprectensfulgnesus Pocock 1902) 6 OTSA Et
M Sprectenus ool Puesl 1905) SOOT SAE WOE
M_ Spiroctenus inermis (Purcell, 1903) DDT SAP ie — dese dal ete
MSprocenuslausPucal,1904 6) OTS SAE WOE
8 Sprocenustiohfoot Pucel902) OSA
F Spioctensigncoltewence 1957 6 UOT SAE KN T
A
B Spiectenusmareytiewt.1919 6 ODS KING
© [SprocteusminorHewit 7913) 8) DOSES
MSprecenuspalidpesPucel| 9086S WOE
J | Spiroctenus pardalina (Simon, 1903) 6 DDT SAE aria
J Sproctenuspestnger(Smon 1905) 6 UOT SAE WOE
F SprecenuspiosusTuser 178 TSE
F Sprecenuspunciustenit 1916 SOOT SENET
Mi SprecenuspucelTucer 176) OTSA WOE
€ Sprocenussapitarus(Pueel, 1902) 6) OSES WOE
F Sprectenus sheen Pure 1903) 4) UOT SAE
M Sprectens ticeartus Puoeh 1909) SOOT SAE WOE
B | Spiroctenus validus (Purcell, 1902) [5] op | SAE WCE rift
FAMILY CAPONIIDAE Simon, 1890
M Le
8 Caprio capensisPucell 906 Siar =isim
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
— | = | — | —
=— | = | = | — | —
236
Ansie S. Dippenaar-Schoeman et al.: South African spiders
wlaolanl|onl\|ol\/ol/n|wol\|o\/ol\/zlze
B
B
'D [CoNSTATUS| ENOEMIGTY PROV |EO|FS|6/KAN| LM) NC [NW | We
Copoiachaiatesoen 1986 ——SOSCS~*~S~SsYCi‘ TS
CepenisefofewPucel 1904S) OTE
CapenisresttoaPuceh soe | SE
CopoishoroveePuceh 1904 A) OTE
Coponinertnas(oPcanbicee a7) 1)
Capon seunde Pocock 008 tT
CaponissimoniPucs. 19048) ORE we
a
Dploorearaeredied 252) ME
DplogerecepenisPucel 904 S| OEE
Dpledenedppewune Hosted 25 EN Ew
Dplogerekorocetedied 15 2) THE
Dplogeromirtawece 19 2) SHE
Dplogeneproniatadées 2015) REE
FAMILY CHEIRACANTHIIDAE Wagner, 1887
walol|ol\/zl/z\/oa\oa\/ol\/ol/zi/oal/zi/zZzl/z/a\/a\/alz/aol/aol(am(amlolwlezl/ai/m ol lolol anianlioal/ol|o
BSS SSS
r-
5
wn
>
m
-
;
Yn
4
i
m
Mm
m
creeconhumseveumsimenees ial we) | id) av [ala] |
Cheecoumaticanmiesenieat A)
Gteaconhumangoenssiowzon7 SSS)
Gteacoum dppeeneton.2007 SS)
Gteecuumfowaioa.205 S| OTe
Cteaconum esearch SA
GteecoummistoeeLon. 207 ES
Geach schneCaporacon
Greece stiwerenisio7007 A) OTE
Gteaconhumesonilemeee 995 SSSA) AS
GteemionssemeniGawenes 2) STEN
Cheemonsamaronsiom.2002
Gteremonemseloa.mme SS)
Gteemonebeven o.2015 SSS
henna boshndeis.omovs ——~SCSCS*~si«d|Cta awe) fe) ||
hears caigeaSiron es Sia | tee | |e) |
team coli lawens wa) ———S—i af we |e Pt} P|
Gheeniraaebewitaw.aons ————SS~s | Ce) ||)
heanionaeunfotsion.zom——SS~dA| te) |)
cheers oes (Svon 9) ———SS~s| tee |) |)
cheonione teedonot. 208 ————S~— af we |e | | fe] |
Ghernione fons ot.2002———S~*~s SSCs
Cheiramiona haddadi Lotz, 2015 za
Cheiramiona hlathikulu Lotz, 2015
a~
N
Zz
m
~)
=)
=
Yn
>
m
aw
N
Zz
m
Cheiramiona hogsbackensis Lotz, 2015
—
Cheiramiona jakobsbaaiensis Lotz, 2015
3
i)
pa
wn
5
m
m m
m)m;m
Cheiramiona kentaniensis Lotz, 2003
Cheiramiona kirkspriggsi Lotz, 2015
-
:
Yn
>
m
aw
N
Zz
m
1 yal
1)
Cheiramiona krugerensis Lotz, 2003
Cheiramiona lajuma Lotz, 2003
-
;
ep)
5
m
Cheiramiona langi Lotz, 2003
~)
=)
=
Yn
>
m
m
QO
m
Cheiramiona lindae Lotz, 2015 iri
Cheiramiona mkhambathi Lotz, 2015 ll
1
Cheiramiona mlawula Lotz, 2003
— —
—
Cheiramiona paradisus Lotz, 2003
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 237
Ansie S. Dippenaar-Schoeman et al.: South African spiders
_ [constaTus eNDeWETY| PROV eo] FS] 6 AN L[M| NE [WW] wo
Grevomovepbabaeaslouais——SOSCS~*~sSSC“‘i AES
Oretomonegschaeliiow.2005 8) wT SAE
Creromonoresiow2009 SAE
Cheramoneroteeto.2015 8) wT)
Oreiomonasmnensstoz 258A SKN
[chetemionatembensistomayis| fe | pon | sae une TT Ty |
[GhetemionaupperbyensibLotwanis S| por | see | | | |
FAMILY CLUBIONIDAE Wagner, 1887
B
B
B
B
B
B
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
Cution aboenassvona i906 ——SSSSS=«dT tS] Yt) faa] |
Citionticamlesentset ttt tt
Cutionssmitgentecwen i929 tT
Cubionaoppoa simon 9102) rH
Cubionbensiteowen 923 8] Ea
Cubion becvetasinon 1897 | TET
Cubioncopenoesimon 1897 | TRE
Cubion ccolortawence 1952 | TE
CubioneduvaraRoowers9s| 8) ES
Cuhiowsodtentessent tf
Cubioneve simon ts? | worm
ea
Cubiowiowencetoovent951 2) tHE
Cubiontinpde Son 1857 | worm
Cubionaetatee Simon 87ST KN
Cubion oteens Simon 7 S| mor
Cubionapongaonactaweren 19528) AE
Cubionpupoistowence 19998] AE
Cubionareiod essen r956 2) tHE
Cubion umpire awenee 19528 | mora
Cubion sitalawence 1952 af
ape,
—
Res WB
xy
5
wn
>
m
Nn A
N | N
m)|)m
Austrophaea zebra Lawrence, 1952 fa; uc | sae
Combale ppenearaeHodéod 20%
‘Cambaliauipes Simon 7856
Coenptehus mutica Wedded 200) TO
Coenoptychus tropicalis (Haddad, 2004)
238
Ansie S. Dippenaar-Schoeman et al.: South African spiders
' [conSTATUS| eNOEMITY PROV |EC|FS|6|KAN| LM) NC [NW We
CopafvopumesaSimen Tees —SS~S A
Copotaiedies 13a
Copvetsercaresind 218 2] tHE
Copuetobousns Suen) A
Copuetobuitewioi 2019S tt
Copuetamagnateed 2018]
CopvetamapuaHesind ors 2] tHE
Cormonmstowercetaaics206 St
Comvonmasenigoum(simon 7699)
Eenornaalnss Howie 2012 SSA ttt
norsinitsHadéos 2002S ENE
Gepieganiossmont9 it]
HorpesasuisepeeBosodeesBocwe 20002) STH
Hones atte Bosseees 8 ocwe 20008) RENE
Honpescoccraus Bosseees&oowe2000 8) SRE
Hones combena Besseaes torque, 2000 S| Roe SAE
Hones rowel Bosses &ooqé 70004) OT SAE
Hones nohuake Bosses ora, 2000 6) oT AEE
Hones Bosse & cae 20006) OT SAE KN
Honpestenopton Bosseeers acme 2000 6 | oT SAE ME
Honpestyteaetosseersateiouts56 | ODEN
Hones mareltossdaers soca 2000 S| UAE KING T
Hones mesemivmu Bosseaes& Jooqe, 2000 §| OT SRE ET
Hones rohorumBosedeerstlecwu 2006) OTE KN
HonpesschoananeeBosedbers inca 2000 2) THE
Hones winmerersiBossees&oomé2000 5) OREN
Mednessasemiavoniowssmoni910 tt
Merenusatenessen ts SME
Mecussmonitessents SSS
Messqusmeninismentse
MessapusmerdenalsHadéedaMbe 2015 | oor RE KN
Messapusrtela Pocock 656) 2) THE
Prmophocsnaaicasinenteo7 SAE
Prnopacsprometessent9) SAE
F Prnophoea ave essen t929) 8) wor
€ Verdopawo lume odiad 2009) SAE
FAMILY CTENIDAE Keyserling, 1877
ee
cers congerFoP-carbnig 658
B Creme puchneni(SimenT60)
elem spectaisLesee 1920
"
FAMYLY CYATHOLIPIDAE Simon, 1894
B
F | Cyatholipus hirsutissimus Simon, 1894 4
F | Cyatholipus icubatus Griswold, 1987
B | Cyatholipus isolatus Griswold, 1987 4
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 239
a a~
N N
Zz Zz
m m
Ansie S. Dippenaar-Schoeman et al.: South African spiders
' [conSTATUS| eNOEMIITY PROV |EC|FS|6/KAN| LM) NC [NW | We
S [Gatotnusqiedinecdaussmon 18 ~~S~; aC SAE
M Ojatolpustrts owed 1967 6) Rae SAE MNES
€soaconusols Grows 1867 SR AEE
F icoshoveqes rowel 1967 6 OSES
M oookyenecieweld 1987S SAE
€ eabureve rowed 1967 6) OEE |
M eabuzwveniscinol 1997 SAE MN
€ Potempsdenipessinon 1694 SOD SAE OE
€ Unacilcpense(Grewols 987) AE
€ UwemiedericusaGiowes 1967 SAE
€ Uwemiueouenuecrewol1987 SAE
€ Uweniuepubhecrowold 1987) SAE
FAMILY CYRTAUCHENIIDAE Simon, 1889
Arete tateronivenrsis) atm | sme || | | | aia] |
Aree menconis(Heniu1919) 8) AE
Arete menbabis (enn 7s16) 8) SAE
Area meyeHentrs1s) 6) OREN
Arey tucoos(Hewt 191) 6) ORE
Arelnpacolonve Pook 1902) | OTE
AvelrypacomuaPwesh 19048) OES
Aner cente Pues. 1909) 6) OT SAE NOE
Arelpeceye(Henwrsrs) 8) OTE
rere dongaaPuce 82) THE
Aree faveotisua Henin 7515) | oT REET
Aree beats Pucel 902) 8) TRE
Areerype magna Hew 7914) 8) OT RE KN
Aree nameqenss Puce 808) S| AEN
Arelnpeniices Pues 1802) 8) AE
Areerpenuéotiewe 1) 8) AE
reste mdes Hew 1923) | wor
Areslipecreiucel 1902) 8) mor
Ares plidpes(Pwee, 190) S| wor RE we
Aresliypepane iene 16) 8) mor
Aresleiypepronee(Hentl 13) AE
reli puolls Puce 90s) | AEN
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Ancylotrypa tookei (Hewitt, 1919)
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M_ Ancylotrypa zuluensis (Lawrence, 1937)
FAMILY DEINOPIDAE C.L. Koch, 1850
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Deinopis cornigera Gerstacker, 1873 Pas | eer
DebopisorndicaPocock 1098
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 240
Ansie S. Dippenaar-Schoeman et al.: South African spiders
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FAMILY DESIDAE Pocock, 1895
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FAMILY DICTYNIDAE O. Pickard-Cambridge, 1871
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g Artec uovaGrowoW eMeRleGravols 1957 3/1 SAE oe
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Izithunzi zondii Labarque, Pérez-Gonzdlez & Griswold, 2017
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, AE
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B | Dysdera crocata C.L. Koch, 1838
FAMILY ENTYPESIDAE Bond, Opatova & Hedin, 2020
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B | Hermacha nigrispinosa Tucker, 1917 5 L SAE E eS ee 1
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B Lepthecus dppereore ios Tamye tye, 2020 S| ODS CET
B | Lepthercus dregei Purcell, 1902 5 jPesites | reac. seceaes i ae
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 241
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘ [GON sTaTus ENORMITY PROV [EC[FS| 6 [KN] Lm] NC [WW] wo
FAMILY ERESIDAE C.L. Koch, 1845
Dressers anqustcps Pucel,1904 SSS | TSCA Swe! | | | |) | |
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Dresses schehesTucer 1920 ASAE
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Paradonea parva (Tucker,1920) Paradonea parva (Tucker,1920) (Tucker, 1920) | STHE a eA am A
Paradonea presleyi Miller, Griswold, Scharff, Rezac, Szuts & me
Marhabaie, 2012
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FAMILY EUAGRIDAE Raven, 1979
B Allothele australis (Purcell, 1903)
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ne
‘Austacheos bag Hadley, Bosssoers Ramen za09 [al uc | sme | | ||| (aii) |
a an et
‘Austrachelas incertus Lawrence,1988 incertus Lawrence, 1938 pe ee ae
Austrachelas pondoensis Haddad, Lyle, Bosselaers & Ramirez,
2009
Austrachelas reavelli Haddad, Lyle, Bosselaers & Ramirez, 2009
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
242
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘> [CON STATUS] ENDEWIGTTY PROV EC|FS|G) KZN LM) NC | NW We
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Drssodela favo Mbo GHodéed2019 SAE |
Drssodelagutate Moo oddad 20198 SAE ET
Drssodela i Mbo& Hadind.2019 6 SAE KING |
DrssodelamacultaMbo GHodéed 2019S) GSAS ETT
DrssodelameoneTucken 9288) |
sfc [ee fee) | tt
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Drssodelaqunqulbecule Tuckey 19238 SAS WE
DrssodelasalcunHewit 1916S) SAE ES
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DrssodelatenebosoLowrence 828 SEN SAE KNE, |
DrssodelatieniMbo&Hadied 2019S OSA ETT
DrssodelatansverseMbo&Hodéod 20198) ODA ME |
Drssodeltineta bo &HadGod 20196 OT SAE ETT
B DrassodefavasinaTucker 1928 S| SAE WOE
B DrassodelaverdaMbo&Hadéad 2019 S| SAE
FAMYLY GNAPHOSIDAE Banks, 1892
® Ammoceus mphaldesDippenaar&Meyec 980 |8|e|S| [tt ttt)
B Ammoxenscoconeussimon tess | STE
B Ammocenus deeds Dppenear Meyer 19808) BD SAE
F AmmocenstabharcusBenok 19722) STE |
B Ammoxenspenterisimen 1896S TT
B Ammoxenspammodomussimon TsO) STE tt
B AmusiacatoraceTusen 9232] SME tt
F AnplsebahearaTwcke, 1923 S| OTSA WE
F meplsafocesTuckent925 A SAE
F mreplessinterogotonisTucken 1923 SSE
F lmeplesarigeTucen 923] use| | |
M Aneplase primers Tsk. 18238] OT SEW
F AnplsascuptuateTucke. 19238 UOT SAE WES
M ApbontouexustalsSimon, 1893S EOE
M ApfataulxinomataTucke,1923 STE
F Aphotalxsigniols Tucker 1928S
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=— | — | — | —
M | Asemesthes modestus Dalmas, 1921 6
B | Asemesthes montanus Tucker, 1923 3] ic | sae
F | Asemesthes nigristernus Dalmas, 1921 fe; pot | sae |
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
Lat
|
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genesis bovitausPuce 1908 —SSSSS« |THE
F conestesdlesTudertsz3 SSS) TET
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F nests decoousPucdh 86 ||
F ress onpesPucs 1900 SSS)
M Aoerestesfodne Tuer 928 SSS~d SSC
F eresteslonteriTuce, 923 SSS*~SS)SCT
F (conestesineousPued.t90@ SS
Aaemestes modes Dales 8 5 a
Asumesties montnusTuckes192 8 a
Aaemestes ngrsenusDaines 926 (T ee a a
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— | —
243
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘D [consTaTUS| eNOEMIGTY PROV EC|FS|6| KAN] LM) NC [NW We
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F AcemeatespaliausPuedl 1900 8A
F AcemeatespamenTuaert2 SAE
F AcemeatespucelTwke.1929 SME
G AcemeatesrfousTukes1923 8 SAE
€ Acemeatessububiussimon T8672 SME
8 caniinaatoboc (sera 907)
€ Canin dpbge tudes SAE
€ cantina copes PacksMuphy 9878S
8 cantina comiea(Tulent) A
€ Caniinanow Penick eMupny. 97 A
8 caniinporesi mont)
8 caniinaproaneuodit906) SME
6 caniiasawseTucle 192A
F DipracuslepoiPucel 19078 SAE
F Dresses bechuncws Tudor T923 2) THE
M Dresses cateranusPucel 19076) OT SAE EST
F DressodscaeausPucdl ts 6 OTSA OE
F Dresses cregePuoa.t907 6) OTE ET
6 trassodesereorPucel 9078 SAE
F Dressodsgootd Puce. 1507 6 OTSA
M DrassodeshaenePucel 9078 SAE
6 DressodslphognausPuceh 1907 8 SAE
F Dressodestraus Puce 19076 SAE
M DrassodesmascuusTuckes 1923) THE
€ Dresses sesqudenusPuceh 68
F DressodssotoiuePucel 907 SME
G DrassodsoplendersTucke 1923S
M DrssodesstaonisTucennszs SAE
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F DrassodstonwaueTuckes1929 6) OTE NE
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LeplodassusbergersisTucke, 19236)
LepocassusfeenosusDaimas 1919) OTSA WOE
Marinarozelotes jaxartensis (Kroneberg, 1875) Cn ea me ae 1
Megarymackon schenenTuxke, 19282) THE
Megamymackon ansveanseTucke, 1928) OARS TTT
MegamymeetionvebxSimen 18872) SE
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McarebeauoneTwte 1923 tA TTT
McarabiptculeBooyoentHedsedz077 | tT
Micaria chrysis (Simon, 1910) Pag Eee le Re Sa etal apf | I 1
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 244
Ansie S. Dippenaar-Schoeman et al.: South African spiders
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cara dubaraoowenaveaed az ———~S~sYSC
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Mearakonnoss Bowen tHoaiad az 8D
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carole Boysen tHodead 21 | ET
Mearamedospra Boysen Heated 277 | DRE
MearaquravenecuoseooysenaHeaiad az) 2) THE
Meorasonbss owen Hoag anzr 6) ODE NE
Meare sult Booyen &Hodded2027 | ODE NET
Mearatesssmesimon i910 | OT SAE
Nombioassats0omes 182 6) OT SAE we
Nombio teres Puce 1506) | wT
Nombienetedaines 19% | TREE
NombiovorowoleaDames 152 a] Et
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NombioveieCuclr 82) 2) SME
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FecclochosinoueTucke 1238) tS
Posocheauis(ucle 28) | tT
Fastolus deseo Heddod 20082)
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FastolustorbaPanickacrintss0 2) THE
Scoophacwsmoioitictes1923 8) AE
Seotopaeusrtaersstowence 19982) THE
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SeoophaeteegausPucd 1907 2) SME
Smopnsbomiuckesva)
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Swophboubtis(Smente) wa
Smoniacopenssvolnes 19208) rE
Smionalesipes Puce 1908) THE
Tepopode apneic 925) AE
Tepopodebamenta(ucer 925) 8) wor
Trephopoda hanoveria Tucker, 1923 T A 1
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YeopheewsawaroumPucel 1907S) SME
YeopheewsbeaweTucer97 S| SE
Neopheewsbpadauetulgen 110
NeopheescopensPucel,1907 SSSA
Xerophaeus capensis Purcell, 1907 SAE
—
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 245
Ansie S. Dippenaar-Schoeman et al.: South African spiders
’ /SowSTATUS|ENBEWTY| PROV o]FS|G| aN] LNW] ws
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a [roorhansrateueruceh wt? ———S—~s | we | we af ef | | |
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a mtieuetnani ————SSSSS—ia| we | sme | | a) |
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a [zoe ate ace vory ———SSSS~w TY te | me | PP
p [zoe davari(onen ion ————SSSSS~d we | |p Pt
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B | Zelotes gooldi (Purcell, 1907) STHE DT oa 1
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B | Zelotes lotzi FitzPatrick, 2007
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 246
M | Zelotes muizenbergensis FitzPatrick, 2007
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Ansie S. Dippenaar-Schoeman et al.: South African spiders
B
B
B
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M
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B
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B
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F
B
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TeoesranbenaeFuPon207——~—~S~iYCi
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ZaloesngonenseFaPavck 2007 | RENE
ZloesotoFipouck 2007 2] tm
Zloesovnberistawencet7 2] tHE
Zaloespatdpestnken1923 2] tHE
Zeloespucripes Puce 1908) | DREN
ZltesquabegersisFPavick 7078) OTE
Zloesredaustawene 1528 2] HE
losreancus(Pvesh 1907) 2) HE
Dlotsresouionriapowck 2007 RE
Doloessoweituces'923 2) tHE
Zeloes sowaus(OPtambrgs 197) tf
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F
eee (Het ABM 5) [5 So ke Se EE Se
faetiietotathratesimen nese, ee | ome ae oa a
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Lae ee nn ee Sn eee
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FAMILY HERSILIIDAE Thorell, 1869
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FAMILY IDIOPIDAE Simon, 1889
M
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F
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B
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
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Ansie S. Dippenaar-Schoeman et al.: South African spiders
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MopscoserewsHenwtsts At
Mopscudenthewints) ST
opsfoves(Pocock 1907) SSS
MopsiniPucewsts) SS tt
Mopsoehoninents STE
Mopsocipes(Hewtut519) SST
Mopsornds Hewints15) SSS OTN
Mopsounneitiew 1913S TT
opshomito Pecos 1902) SSE
Hopshepunitewet919) THE
Mopshioms(hewn ts) SSS
opskenariows Puce 190) SS
opsmicops Henin ts13) BOT
Mopsmenicobtewtsi916 SSS tt
Mopsmessanbios Hew) tHE
Mopsrigepiosustenirsis)
opscoreaus Pook 190) SSS OTE
MopspostewintsiS SOT
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Mopepuustutenton7 At
Wopeswvears Hew 1925) SOT
Wopettee OP-Cambiage 67ST
Mpevendori(iew.1925) SE
Sepegarabatami(iewint9i3) SOE
Swpegarapeuipnuos(Hewtt55) St
swpegaratenondlrse(ewwrss) 8 tA
FAMILY ISCHNOTHELIDAE F. O. Pickard-Cambridge, 1897
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FAMILY LINYPHIIDAE Blackwall, 1859
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
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weinuspuusoP-cambrige 1904 6S
Meonidsdenbopssmen 164 2) tHE
Memessistadeoum(Gerondt92) ta
etolpuphanesaniars oowe.r904 8) SE
Metlepharespeexows Son fogs 1821) tA
Merton stems @aves.1085) 1) tt ttt
Norevefanmeavanesdngen 19654) OT SAE
NereneratlersiovanHesdngen 196 8] SAE
Notescpisaustalssimon 1696) OTE
Oseoiusmelrepous(OP-Conbides 1679) 0) Tt
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g Andommeratfaysmon 59 a] 0] swe | a | fa | | | |
M CoysophuscheascasSimon 3to SOT SAE OE
M Coyssphuspacvaussinon 198 SOT SAE OE
F oyssphusuicabrsinon 19036 OT SAE OE
F Aracbcedsdenaustawercs 2 2) THE
F Aracbocessewesvissinen tes? 6) OTE
M AracbosessevoTueerrs0 8) ET
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F AtueboctesounolausTucke.192 2) THE
FAMILY LYCOSIDAE Sundevall, 1833
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 249
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 250
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘D [ConSTATUS| =NOEWIITY PROV |Eo|FS|6| KAN] LM) NC [NW | We
G [PawossaasspopsPucds# —S~S~—~ SSIES
F PaosaenueaaRoever 19596) OTSA
€ Paaosahivande(OP-Combidgs 1679)
€ Pacosolpomrucdits SME
F PaaosayessinePucel 1903 OTSA
€ Paaosamanubiaasinon 1898 SME
€ Paaosanosvoum Alewehelcésoowe 7992 2) THE
€ PaaosascnenenPucsl 19032) wT
€ PaaosoumloPuedwts08
F Passer auter(Smon 188) 6) OTE
F Plas aticone oewent960) 2) mE |
6 Provppaaivens(Smon 188) SME
€ Proevppa dial Puce 190) SME TT
a ae ae Bo ESI EE
® Poevepahirsua Ruseismih 198) |THE
® Poevpralihfotipucel 1908 SAE
B Poevpraschranei(Pucel 1905) STE TT
M Poevppawanessi(usseSmith 969) 8) SAE
B Peraroraarbusoub Pucel908) 8) SAE
B | Pterartoria caldaria Purcell, 1903 (43) recor a) Se ae 1
G Pleraroracedrbagersisussel-Smith@Robens 2017 §| eR SAE WOES
® PleraroriaconfuseRusselsmiienobens,2017 4) SAE
F PleratorafsvitoePucel, 19036) OTSA ET
G PlerarorafovlmtaPucel1903 S| DSA WOE
B PraroroltnittsPucel 03 ASAE
® PerarorapoystctaPucel 903 S| SAE WOE
F erarorasgoe(Pucel, 1903) | OTST
B Prerararasubeworea (Puce 1909) ASAE
B Schzocosa daring Pocock 1858) CSTE TT
B sohzocosasubpesoata(Sinon 1910) 2ST
B Tabeshetocuatesvend 19S
M TiobearataenisRussebsmit, 1962 2ST TT
B TbesrigneepsPuceh 1903 SEE
B |Trabea ornatipalpis Russellsmith, 19828) SAE tt ttt
B |TrabeapurceliRoewen 1951 tt) ttt ttt
B Trabearubriceps Lawrence, 19522) STHE ttt
M wee] [Tt ttt ft
B |Trabea varia Purcell 1908 8S Et
B Ticassadeserticola simon 19102) STHE tt
F | Trochosa albipilosa (Roewer, 1960) 6) OT SAE ECE
F | Trochosa eugeni (Roewer 1951) f6| por | SAE | NCE iWon
F |TrochosamodestaRoewer, 1960 | 6) OT SAE PE
F |TrochosanigerrimaRoewer, 1960 | 6) OT SAE PE
B Wedleosaonckatawence 1927 A
B Zenonneabocawitetawence 1962 8) ST TT
F |Zenoninamystacina Simon,1898 2) tC STHE | ttt tt
FAMILY MICROSTIGMATIDAE Roewer, 1942
B | Microstigmata amatola Griswold, 1985
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 251
Ansie S. Dippenaar-Schoeman et al.: South African spiders
Se a 6
F MocsionatalawercdGiowoKs1985 OTT
© Mowsionselngpestawerce 199 SSS
€ Mocslonaeutionta Gaon 985
€ Mocsionascubense(lawene 99)
FAMILY MIGIDAE Simon, 1889
F Meogvesabineaioarewines «| wor | we we | | | | ial | |
F MeomivesamputaGiowe 1987 —SSSS~CSS YS
F Meomivesboeiennisis 8 or
F MecgnvescuaenHewnrsis 8) OTE
F Meomives yn oP-Conbasete75 SSSA)
€ Meogvesimemedatewt 798) Rae AE WOE
F MeggeslepaldiPucel 150 6) OTE WOE
F MeoqivestosireGrswoli1867 SSN EWC
F MeooveamacnsHonntsis ET
F MeoovesmordrPucsl i909 SSS) EWE
F Mecmvespowionnonenis 8)
8 Meoovespeneuaisment9 8)
B 1
B
B
B
B
AE
es
Mosidgespyminewit.tot4 2) SE
Moggrdgesquercnasimen 98S EN SAE WOE
Moguidgearpcoanents SAE
MogaiigearpcbdesHewit18 8) DOT SAE EE
Moggridgea teresae Griswold, 1987 WCE
Moggridgea terrestris Hewitt, 1914
Moggridgea terricola Simon, 1903
B | Poecilomigas abrahami (0.P.-Cambridge, 1889)
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M | Poecilomigas elegans Griswold, 1987
FAMILY MIMETIDAE Simon, 1881
B | Anansi natalensis (Lawrence, 1938)
F | Ero capensis Simon, 1895
B | Ero lawrencei Unzicker, 1966
F | Mimetus cornutus Lawrence, 1947
FAMILY MITURGIDAE Simon, 1886
M | Parapostenus hewitt Lessert, 1923 sf te | se Tt tt a
F_Voraptusafris Lessert, 1925 pS oe See ieee
FAMILY MYSMENIDAE Petrunkevitch, 1928
Uroctea septemnotata Tucker, 1920 STHE
Uroecobius ecribellatus Kullmann & Zimmermann, 1976 rs| ue | sae Ef dist od ltl ledl 1
Uroctea schinzi Simon, 1887 STHE ||
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
GON STATUS ENDEWITY| PROV [EC[FS|6 KN] LM] NC [Ww] WC
—y
252
Ansie S. Dippenaar-Schoeman et al.: South African spiders
' [consTaTUS| eNOEWIITY PROV |EC|FS|6| KAN] |W) NC [NW | We
FAMILY OONOPIDAE Simon, 1890
Ausiolooopsganuausiiown ins 8] | swe | aa) | | | |
Ausilonopshadied Pick Owing 202) THE
AusilonopsshePonikOweng 20714 SAE
Dares codeeisebsaune 2 | tsa
Dame grovel seated. 20126 OD SAE
Dyedenecapense Simon 19076) OD SAE OE
Dyed specu Son 1972)
Ganwsononroamsroishewntis 8] SAE
Ganasonaptohimicoblowenes 9478)
Ganasomaphe lies Lawence 19526) oT aE KING t
Conmpscscastent 9752S
Cpopecameticasinont093 At
Opopeespcsatlawerce 195) 1 tt
Gres orcasimonts3 | OTE
Creetnafomesewadaoowé 222) THE
Pseutescophila parse Simon 19076) OTe we
F Telos arovasloveSimon 19076) OTE
FAMILY ORSOLOBIDAE Cooke, 1965
FlatlousassleGiowséePimick 967 =| 8| vor | sae wee! | | | | | | | |i
M AflouscapessGionodePowick 1987 SAE
€ Azaiaobuslowencd Growolda Planck 1967 2) STE
F cabwusdedoreued 06SEC
FAMILY OXYOPIDAE Thorell, 1869
Hanae tonconistesonn vay tf te] we | (a | a dais) |
Honaaotacanatitessen 9919)
Hanatava rotors (awencs 128) 2) SMHS
Honaabvonsocaigaasinon 10981)
Manat stondessen 7929) 2] tHE
Oyopesafinstesee tS
Onyopesanguiarstesee tS
Cr
Onyopesbomevieeeet 19992]
Oyopesdotitessentns
Onyopescasonsstawenen 17 2] tHE
Onyopesctopiesenrsz7 tt
Ones oenensteee wen
Oxyopes dumonti (Vinson, 1863) ya
Oxyopes falconeri Lessert, 1915 1
Oxyopes flavipalpis (Lucas, 1858)
Oxyopes galla Caporiacco, 1941
Oxyopes hoggi Lessert, 1915
Oxyopes jacksoni Lessert, 1915 a
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Oxyopes longispinosus Lawrence, 1938 3 | SA
Oxyopes pallidecoloratus Strand, 1906
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Oxyopes personatus Simon, 1896 6 |
Oxyopes russoi Caporiacco, 1940
Oxyopes schenkeli Lessert, 1917
Oxyopes singularis Lessert, 1927
Oxyopes sjostedti Lessert, 1915
SA
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 253
Ansie S. Dippenaar-Schoeman et al.: South African spiders
' [conSTaTUS| eNOEMIGTY PROV EC|FS|6| KAN] LM) NC [NW We
F OnopessidebeineCoporaccovoh ——SSCS~—SSSCi
G Onopesbecuaustesentts tw |
B Onopesmonaustesents tw
M Oopesvandensdtessen tos
€ Oopesvepceangeiteowe 1946 A
€ Pavcelacwatea twee 187 SME
G Pewctatuasivnson tes it
g Pewctamecuoeroreck 900 SME
€ Pevcetamadseae Von Neko DppeoarScheenan 1994/2) UC STHE TT
€ PevcetaniceeVanNeler&DoerserSchoonontse4 3c SAE
€ Pevcetpucra Bochva 665 tw
G Pauctasvinakaccn te tw
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FAMILY PALPIMANIDAE Thorell, 1870
g Dephoocetusbipasiaussinon ies 2] uc | ome (|| a) | [a iafa | [a
€ PabimanisamausPoosck 10988) AE
J Papmanseweustawencet927 2) wor SME
G PabianscapenisSimon tess SAE
M Pabimanuscrdeiteswn 996 OT SAE
M Pabinanisgiuaytesen rie 2) THE
M Pabimaniegobutersimon 10938) OTE
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Paimarusranequensssion 9102) THE
Papmaruspuccuussimon 15106) OT SRE NE
Papmaruspotentamens 957 8) AE
Papmaruspsewomaustawence 19523) SAE
Papmarussubamaustawencs 1987 6) oT RE KN
F Pabimanssarsvniassimen 1893) SAE
M Pabimansttecusustawerce 19526) bor SAE MNT
DDT SAE 1
[Penestomus eroeseriDippenaarschoeman,1989 | ¢|_— br | sae | ce | 1] | | | | | | |
[Penestomus egaziniMille, Griswold aHaddad,2010 | 8| Rae || sae | ce | 1] | | | | | | |
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F Mra varigote (Son 1895)
F Phiodonus bgbbus usta Lawence 1328
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 254
Fa Bs FS Sa SS BS
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘> [CON STATUS] ENDEMIGTTY PROV EC|FS|G/ KZN LM) NC | NW We
B Phicromsguneensomilow9@9 AT
F Phiaromusporiustessen 1919
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F Phioaromus wip Simon 1906 OT SAE OE
B Svemuspunctostawence "968 2] SME TT
F Tanatsaticanuskarch 167 t)A
G ThanatsatoniowsBefond.1996 tA
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® Thanatusiobici(@udouinte2s) |
G Thanatsiomotel zeae 966 t]
F ThanatsnamaquensisSimon 19108)
G Thanatswiarssimente7 |
G Tbelusamaustessen9@8 tA
F Metis austaie(Simon 190) |S
F Telus buneiarib awence 9822) SHE
M Tiel cobus Vanden Berg ippenaarSehoeman 19941) AE
B ibelusdemangelseceaue 1964
Bets evipesCaporicco19@8 |
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B Thetis holdoylawence 1962 AT tt
G Tbeluskboncensbtese. 1919
B Thetis minortessen 99 AT
G Tbelus serepunctaussimen 907
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FAMILY PHOLCIDAE C. L. Koch, 1850
‘AtereateniaWalckewe, 1637] | | |) it
Grossoprzeoni@lecwal 867) |
Leptopholcus gracilis Berland, 1920 a ai oe
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Quamtaaciite(Lawence 1998) 8A
Quamtansembuenitbe.z003 8) SAE
Quomtangertabenibe. 20038) eA
Quomtanafimenuber 208 SAE
Quamtanshecontube, 208 8 AT
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Quamtana vidal Huber, 2003
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 255
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘ [Gow sTaTus ENDEWCITY PROV [EC[FS| 6 aN] Lm] NC [Ww
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Srwrngopis debe Ween 20026) ME
SeingopusmivaneHbec 2022] Re THE
SeingopusratalersioLawerce 19472] THE tT
Seingopusndumo ber 2072 | Rae
Srwingopissedobegtibew2002 AE
SreingopuswbTibe.2012— a] AE
Spemopton gordmeeHube. ino 8) Rae SRE WOE
Spemoptonpenbaee 20058) OTSA
Spumopton peninsubelawercs 1866 S| Roe SAE OE
€ Spemophorschoemarsetue2003 6) OR SAE wee |
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FAMILY PHYXELIDIDAE Lehtinen, 1967
Lamatedsinaacrswow1550 «| oor | swe twee! | | | | | | |
Molo delcauie iow 7990 8) OEE
Moloto ropes Puce 08) | Rew
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Nanaquecmehetenate Pocock 190) | 0D SRE NOE
NanaqurcineMoioareGioweld1990 6) 0D AEN
Namaqurcettounauie iow 1990 S| ODEN
NanaqurcinevopstaGiowoK, 19908) AEE
Pryde motapaersieSimon tee AE
Pengolanechyooreie rowel 19908 AE
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
Srwingopisfevsbedhuber22 AE
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256
Ansie S. Dippenaar-Schoeman et al.: South African spiders
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TYovesoormoneeswiowoa 10 ——~S~idYCi
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F hosmepesromeaiense Roenen1965) 2 SHE
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 257
Ansie S. Dippenaar-Schoeman et al.: South African spiders
' [conSTATUS| eNOEMIITY PROV |EC|FS|6/KAN| LM) NC [NW | We
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TreunamecubePwesl 1907 2) STM
TheunamuaPuceh 907 | OTE
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Treunapucsucke 238
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F Treune wuenoslawenes 1947S) OTE NE |
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pw ewe TTT TT
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Aetutuscisttopapussimon 1902] sme || | |] | tl
‘Aatcileatera Wesdiowsa,200) tT
‘AatciletpunctataPecktom 8 Peckham. 1908) 4 OTST
‘Aafia bausi (Peckham &Peckbom.7905) 1) ET
‘Acta cegans (Wesclowske&Cunming 2000) 2 tHE TTT
‘Acta sonica(Simon 1902) OTSA
‘Affe imtotor(Wesclowsks @Hadésd 2073) 5) OSA OT
‘Acie farnoeHodéod&Wesckwska 2017) 6) OTSA ET
‘Ace enustub Wesoowsko& Hades 2005) 4) SAE TT
‘Affe zvenss(Hadéed esoionsa,2019) 6) DAE
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 258
Ansie S. Dippenaar-Schoeman et al.: South African spiders
zininiw
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Cembalea plumosa (Lessert, 1925)
Cembalea triloris Haddad & Wesotowska, 2011
Chinophrys trifasciata Wesotowska, Azarkina & Russell-Smith,
2014
Cyrba boveyi Lessert, 1933
Cyrba dotata Peckham & Peckham, 1903
Cyrba lineata Wanless, 1984
Cyrba nigrimana Simon, 1900
Dendryphantes acutus Wesotowska & Haddad, 2014
Dendryphantes hararensis Wesotowska & Cumming, 2008
Dendryphantes limpopo Wesotowska & Haddad, 2013
Dendryphantes matumi Haddad & Wesotowska, 2013
Dendryphantes neethlingi Haddad & Wesotowska, 2013
Dendryphantes purcelli Peckham & Peckham, 1903
Dendryphantes rafalskii Wesotowska, 2000
Dendryphantes schultzei Simon, 1910
Dendryphantes silvestris Wesotowska & Haddad, 2013
Euophrys bifida Wesotowska, Azarkina & Russell-Smith, 2014
Euophrys capicola Simon, 1901
Euophrys cochlea Wesotowska, Azarkina & Russell-Smith, 2014
Euophrys elizabethae Wesotowska, Azarkina & Russell-Smith, 2014
Euophrys falciger Wesotowska, Azarkina & Russell-Smith, 2014
Euophrys gracilis Wesotowska, Azarkina & Russell-Smith, 2014
Euophrys leipoldti Peckham & Peckham, 1903
Euophrys limpopo Wesotowska, Azarkina & Russell-Smith, 2014
Euophrys menemerella Strand, 1909
Euophrys meridionalis Wesotowska, Azarkina & Russell-Smith,
2014
Euophrys miranda Wesotowska, Azarkina & Russell-Smith 2014
Euophrys nana Wesotowska, Azarkina & Russell-Smith, 2014
Euophrys purcelli Peckham & Peckham, 1903
Euophrys recta Wesotowska, Azarkina & Russell-Smith, 2014
Euophrys subtilis Wesotowska, Azarkina & Russell-Smith, 2014
Euophrys uphami (Peckham & Peckham, 1903)
Evarcha alba (Peckham & Peckham, 1903)
Evarcha acuta Wesotowska, 2006
Evarcha amanzi Wesotowska & Haddad, 2018
Evarcha annae (Peckham & Peckham, 1903)
Evarcha brinki Haddad & Wesotowska, 2011
Evarcha denticulata Haddad & Wesotowska, 2013
Evarcha flagellaris Haddad & Wesotowska, 2011
Evarcha ignea Wesotowska & Cumming, 2008
Evarcha karas Wesotowska, 2011
Evarcha mirabilis Wesotowska & Haddad, 2009
Evarcha prosimilis Wesotowska & Cumming, 2008
Evarcha striolata Wesotowska & Haddad, 2009
Evarcha villosa Wesotowska & Haddad, 2018
Evarcha vittula Haddad & Wesotowska, 2011
Evarcha werneri (Simon, 1906)
Evarcha zimbabwensis Wesotowska & Cumming, 2008
D | CON STATUS | ENDEMICITY | PROV EC FS; G_ KZN
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
STHE
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1
259
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘CON STATUS] ENDEWIGTTY PROV EC|FS|G)K2N|L_M) NC | NW We
FesucuahediodiNatinatFood204 ASAE |
Fesuclaeoyoe ReatnaFoors2014 2) STE
Feswoubrobusus AearnasFoo2or4 8 BDA |
Goleopueloinon ies) tT
obrocestum aiconum Wesolowsha &Woddod2009 8G SAES |
Hobrocestum abimanum simon 19018) SAE OE
obrocestm auicomum WedGode Wessiowsk, 2018 ooT SAE EFT
Hobrocestumfovianumsinon 19018) ODA wOER TT
obrocestum uae Peckham 8Pecktam, 19084 DOT SAE TTT
obrocestum lcuentum Pecktam &Peckhom.1908 4 DOT SAE
obrocestum sapiens Peckham gPecktam 190) 2 STM
Habrocestum superbum Wesotowska, 2000 Pe Je | sree SS ale
obrocestum tarzancum Wesolowska& RuskSmih, 20001) EEA |
Harmochirus luculentus Simon, 1885 a a ee eo 1
osaineledstnte Haddad aiesolowska 20134) SAE |
Hosarusadasoni(udoun.182)
eloptanus aecorensis Wesclowsta,1986 1) MAE
Heloptanusaficonus Wesoiowsha 1986 S| EN SAE EF
HelophonusbelusWescowsta.1986 DOT SAE WOE
eloptanusberondiLawence 1957 | DOT SAE KNE | TT
ZlZzlmal/al(al lalallala l/al/Zzl/zZz\/aol/al/ai in| ala lwalaol/m\a\/a al Zzl/alai/aoini a(n Zz\/m/wala\/wal al al/wal/Zzi/Zz/mn|alZz/zZz\|aola\a\|a/\|o
HetopharustoucseWescowsta, 19962 tHE
HetophauscapenasWescowsta, 19064) tA
Hetoprauscapcabsmon 901 tHE
HetephansctoeiWesobwste 7008 SSS) tA TTT
Heteprascowsersmoni9o1 tA
Hetephausdeomets Peck Peahoni908 2 tHE
Hetephansdebissmont901 tt Tt
Hotephaus densratnsWesdonska 19861)
Hetophauseesercdeimon 1901S tS WET
Helephaus designs PesihamaPeckham 19086) OOT SAE
HelepanisfschausWessowstet906 1) tA
Helephas garieusWessowstaHediod 738A
Helepanistandersimen 86 tt
HelephashasousWesooveka 1986 tHE
HelepanishorerWesdbowska 1986 SSS OTE
Helephaueifausus(Peckhamapeciom 7505) 1, te
HeleparusispeausWessowster966 tt
Holeparistscerivesoovsta 1986 tT
HelepanismachollPettom &Peckbom.1903 | oT AEE
Helephanue mails Wesoiowsta 1966 S| OTE Owe
HeleparismodeusPeckomaPeatent90 1) tA
HelepanisnansWesoovela 2009S
a
Helephansortesesinenvees tt
Helepanispaclaresimon i901 tHE TT
HelepanispaiperWescowsta 1986 Tt
we (wel |T{ T[Pl i
HelephauspstocaeWessowste, 20032)
HelepanisporerosueWesobvete 18868 OTE WE
HelepanisprawiPectam aPeanen.t503 2, tHE
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 260
Ansie S. Dippenaar-Schoeman et al.: South African spiders
walzlzl\al\a\/oal\/oa|/nl|oa\/oal\/oal/ol/z/al/aoi(niol/ol(/ni/ol(an(mlel/olezl/ezl/zZz/o/ao/ mi amlwolzl/al/oal/aol/m an noal|ol/ol|ol\/ol/zl/o\o\n|\oa\|o
‘ |on STATUS| ENDEMICTY| PROV /EC|FS| 6|KN| L|M|NC_ NW Wo
Heliophanus proszynskii Wesotowska, 2003 2 aE Sa line 1
elophans pygmaeus Wescowsta.RusselSmit,2000 1) uC) AE |
Heliophanus redimitus Simon, 1910 fe} pot | sac |nce} | | | | [| fa] |
Heliophanus sororius Wesotowska, 2003 hey ke. | STE | lala) fa,
eloptanustemtophagus Wessiowske@Hedded2002 4M SAE TT
elophon ther Wesolowska 2096) vOT SAE ET
fe ome fe
Heloptonustepiussimon to STE TTT
Hipogeorgus Pecktam Peckham 7682) 1) AE
Holcbeiselewesimon 190A |
Holeooetszuuensstawence 1997 AE
HomelatuscoraceusSimon 1902 AE
Homalatusobscuus Peckham &Peckham.t903 6 DOT SAE WOE
omolatus punctatus Pectham &Peckham.t908 S$) EN SAE KNE, |
Hylusergrotoussmon 1902 AR
HylusbretarisSimon,1902 St) ATP tt
Hylus dots Peckham aPeckhan.t805) AES TT TT
Hylusfavescos simon 19028) DOT SAE NE |
HylusweleveniPeckhomaPeckiams1908 1 AES TTT
Tous dendyphantides Stand, 1909 8) DOT SAE WOE
fous deseronmsimon 19016 SAE WOE
Teusisoldus(Wesclowsta,199) | STE | TTT Tt
Tous nigrcaudusWesclowska Haddad, 20098) EN SAE KNE, |
Tous pubes Haddad Wesclowsia.2011 4) wT) SAE
anatuspincpals Wesclowsha,2000 1 Af
KinaaficanaPecthamaecknom1902 2k SMT
KimaataPecknam ecto 1908 AE |
Kima variabilis Peckham & Peckham, 1903 iS] te | BAB lal ae] fa 1
LangluituscdarbagenisHoddod&Wesoloska,2018 6 «DOT SAE WOE
LangluiuskrugerWesclowska Hadded. 20186) wor SAE |
Langluiusminaus Wesclowska &Curming. 20172) THE |
Langlais amicus Wesotwska.20 2) THE |
Largelurius sqamigerWesclowska &Haddd,2018 6) UDA KIN T
Langone ete Weslowska&Cumming 2012) THE
invomticetonace ta? id] ef ame | PT
Lenore hteaavediedeMeeooweinwovi SS) ea
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nore maniaisiren tant ——_____[«| oor [ae [et] {} ppp
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 261
Ansie S. Dippenaar-Schoeman et al.: South African spiders
D\D\/n\/an/nia/ai/a/al/ain|wDlwDl/Zz/Zz/a\Zz\/al\wmal/Zzl/al/Zz/al/ZzlZzl/ol/Zzim mn wala\wal/wal\Zz/zZzl/al/z(mlZzlaol/aol/ainlolow
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[Menemerus bifureus Wesotowske, 1999 || tc || ty] ft} tia}
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 266
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘» [CON STATUS) ENDEWIGTTY PROV EC|FS|G)KZN|L MNO. NW
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F Anphons reals lawrence 194) 8) OT SAE WOE
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F | Selenops dilon Corronca, 2002 SAE |
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F | Selenops tonteldoos Corronca, 2005 6) DDT SAE
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
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Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘> [CoN STATUS) ENDEMIGTTY PROV EC|FS|G) KZN LM) NC | NW We
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B EupassusschoenanaeMoredmand 20182) SME tT
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B Olosauicomis(Smontee0) |
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Palystes karooensis Croeser, 1996
Palystes kreuzmanni Jager & Kunz, 2010
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Palystes leroyorum Croeser, 1996
Palystes lunatus Pocock, 1896 6 DDT
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 268
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B
B
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Ssusimopus cases Puce. 19036) OT SAE
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
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Ansie S. Dippenaar-Schoeman et al.: South African spiders
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€ Stsiopusspnoaistewt 14S) ODE ET
M Stsimopusctntugensc Hews 156) OT SAE ET
F Susimopusautiscwstewinisis 6) OTE
€ StsiopusysonHewnvtss 6) OSE
6 Stsmopus nspnesus Puce 1903 SSS) OSES
F StsimopusumiatcusPucel ts 8) OTE ET
FAMILY SYMPHYTOGNATHIDAE Hickman, 1931
F |symphyconata mina crowor1967 «6 oor | swe wane! | | 1a | | | |
€ cargoes obse ong eHeded m8 6) ODE ME
FAMILY TELEMIDAE Fage, 1913
per fe we TTT TTT
FAMILY TETRAGNATHIDAE Menge, 1866
b|dipn oor Ometia Mask aivin2az0 [| uc |e | itl ft (alt |
M Dip cyrum Ome, Marusk& Lye 2020 6) DOT SAE ME
Bion simonivous 1950s] ut
F Dip vandewatee Ome Maruska tye, 2020 6) DoT SAE ET
B Dip wesoonstae Omeko Mowsk&tve 2020/8) RARE SAE
B | Glenognatha argyrostilba (O.P-Cambridge, 1876) Oy) SH | ee WD aaa oi
B lewaweanyescensBenot 1978 TT
F tewowgeauonoumsvend 1907 S|
B Lewawedecoata(achwal 186) tt Tt
B levcouefesva Baca 186) 1) TTT
J Lewawpefohotensssvand 908 6A WE
B LewowgehbonterssTulgen ts)
B LewowelevndeiGucunsi 1901) TT
F lewowemedensslessen 1950) tA tt
M LewowertiomeenssKust960 it) tA] tt
FiMetameuenssTulgen 1910 OTT
B
B
B
B
J
B
B
B
B
B
B
B
F
MetelnehaddedMenskelaen 28 SDE
Paclypaalelepiomene 19821
Parlor zope osmenstsossiees, 19941 wT
Tevagnataogteniskgeeing 1085S :
Tevagnatacotfa Seon 905) SSS)
Tevagataceyonea Panis 188 SO)
TeragnatademiseLkoohte2 SSC)
Taragataisns(Smonte) SS
Teragnatajecusertulgen 1990 SSSSC~iSC
Tevagataepeingsmon 9)
Tevagnatantens(tudoun.te26) SS)
Tevagata swbaqometeOuna 1985
evens yi OParbiepe 906 OTSA
G TarapraveuniconsTulgent10 SS
g Tarepratevemiomistmenon tb)
FAMILY THERAPHOSIDAE Thorell, 1869
B | Augacephalus breyeri (Hewitt, 1919)
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 270
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘0 [oowSTaTUS|ENDEWIITY PROV [EC|FS/G KZN LM NC [NW wc
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achionpuspretoraePuceh 904 SOOT SAE
BrachonpusrobutsPocodk 97ST SAE
BrchionpustistisPucel 1903S OTSA
CeatgnusbecheephausHewi 19194) wT SAE
GertogysdeingiPococh 1657S
GertpyspouberiGalon 2005 SSS) SE
Hopestigave(auelletes2) S| sw
HopestbovonePucal tet SAE
Horacio caferane Waldenees 1657) SSS tA
Horace chysogasterPooock 1057 S| MESSER
Horacio cuatrPooock 1656S SEN
Hopes covpesPocock 857 SSA ET
Hopacta detaerPucel 1902 SSS SASH
Hopes ggasPocok 1098S SAE
Horacio nanitoniPocodk 1902S SET
Hopestoiestorecok 1897) TSE
Hapaciotnata GimonT0)6)OOTE
HapactomansiPucek2 | SAE WE
HopestonanaquenssPuceh i902) SA
HapectpuchpesPooock 1901S) SAE
Hopactotonanwseest67 SET
Hopes domicolaPucel, 1905S) WOE
HapacielahelrsePucel 19038) DOTS WE
HapacielakarooiePucel 19028) OTSA WE
HorpacieloidarPucel 1908S OTSA
Hopactelaighfoouucel 9028) DOTS WE
HopactelalongpesPuceh 9028) DOTS WE
HopectelemaprePucel 1905S OTSA ET
HapecteleorewiGalon 208) TSE
HopscteleschworiPucel 908 OTSA ET
HopectolesphosePuwcel 1900S OTSA
HopecteleeeoviPucel 1902S) OTSA WOE
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 271
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Ansie S. Dippenaar-Schoeman et al.: South African spiders
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Proorcidaebuneo(Smon 695) —SSS~—SSYC‘i SS
Prooreida rincatio(Svand1909) —SSS~*~—siSSSCi Tew
Pryosome marae obers,198) ——=SS~—sSCi SS
Pryosome spindon Fobens 1876) SSS
Pavicknamneon @ocerbewesvond 190) SSO)
Rronpheesafinetessents56SSS~| CSET
Rronphoes esia(Smen 1673) —=SS~S~S~—SSC
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Sted ereonfomis (OP-Conbiage 187) ——~=~S~—iO)S
Seated ge Lawence 196) SSSS=S) STE
Seated fosteeDppeearScroomenawilles1992 4, CSE
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 272
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘> [Con STATUS] ENDEWIGTTY PROV EC|FS|G)K2N|L MNO | NW WC
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B Anseructei(lessen9) St] te |e | ttt a [ati ta
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F Borboropacus avsals(Lawence 987) S| DOTS KN
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F DiscorfaniFoge 192 | SMES
F Disa vibes Srond909 TSE
MFmicusaboomis(essen. 1929) 8) SAE
M Fmicusipncitus Caporacco9#1 1) ATT
B cuaesiacongoenss(lessen 1948) 1) A
B HeraeisalenonesiVenNekei&DppenarSchoemen 20138 te SAE TT
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B Heraeuscassspiustawence 92 1) At TT
B HeraeusforVanNeket& DppenoarSchoeman 20188) SAE TT
M Heraeus mulznbeVanNekek& OippenaarSchoeman 20% 6 oR SAE WOE
B Heraeus petewebbiVanNekei&DippenearSchoeman 2018 2) ke SHE TT TT
F Heraeus sossusielVanNiekerk&DppencerSchoeman,2018 2/16 SHE
B | Heriaeus transvaalicus Simon, 1895 fa) ie | eee ee
® Heraeus sanderVan Niekerk&DppenserSchoeman 20181) AE
B Heraeus zoniVen eke & OppeoarSchoemen 20138 SAE TT
F
B
B
B
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B
B
B
B
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B
B
B
B
B
B
B
B
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B
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B
B
Hewttogocistessen 1928) AES Tt
Holopeusabtarissimonte5 AS
olopeus ania DpperaarSchoenan,T986 4) SAE
Meumenopsrabrodecortusmilo. 1942 AES TTT TT
MonaesesaustiusSimon 1910 AT
MonaesesfscusDippenaarschoeman.t966 8) SAE
MonaesesgbbusDippenaarschoemant968 8) SAE TT
MonacsesgrseusPowsit897 A
Monacsesparaiowusuuces 868) tt
MonaesespustlosusPavesiN@85 ART
Monaesesquadubeclaustawenee 19271) ART TT
Mptarefovoqate Lawrence 1952) St) tA
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Mystaria lindaicapensis Lewis & Dippenaar-Schoeman, 2014
Mystaria irmatrix Lewis & Dippenaar-Schoeman, 2014 STHE
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Mystaria occidentalis (Millot, 1942)
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Mystaria variabilis (Lessert, 1919)
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 273
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘P [CON STATUS) ENDEMIGTTY PROV EC|FS|G)KZN|L MNO. NW
B [Onptincoensadeewel965 TA
B Poctactescompactslawene, 1947 8) SAE
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B Pombomismainiesen 1919
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® Pubomisploss DpperserSchoemanFoord2020 2) SHE
B Puesmod qudivberuta ede 966 1A
M Pheenopomarigopuncttum(OP-Cambrdge 1869) 8ST
B PhereydescarnaeDippengarSchoeman 19804 SAE
F PhreydesionaeDippeaarSchoeman,1980 1) AE
B Phreqydeslucne DpperaarSchoeman 980 8| SAE TT
B Paredes ncleas/DppenaarSchoemon,T880 8) SAE
B Phreydes tuberous OPCambrige, tee) STE TT
B Phrecydeszebolawence 27 tT
B Phynaracine melletaoibessen 19882) STHE TT
B Prynarctnengose(aneller@oe) St) A
F PloythomisusdesercoleLawence e362) SME
F Poythombusjbbitowence 19688) SAE
F Playtomisus bys Lawence 19686) OT SAE KE |
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B uncnadepessasimonss0é St) AT
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B Runcnbinecta (LKochte7s) |
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B uncnatopicasimon 1907
B Simoruscapensssimen 1898 A
B | Simorcus cotti Lessert, 1936 i Sr oases
B Simoreushadied Von Nekek&DippenaarSchoeman 20108 NT SAE WOE
B SimoreusioVan Nekerk&DpperaarScheemen.2o0 2) SHE
B Smodiomuscorongersimen 1895) AE
F Siphropelagaciislawence 1962 8ST
B stbhropusofinstessen923 |THE TT TT
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B Stpopusinemeduswi 192 tt
B Slime ndumi ital &ippenoarSchoemanaovt 2) STE] tT
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B Synemadena(audount@2) |
B Synemaimioti«Pavesite69) TT
F SyemalengheiDany 1907 |
G Synemamandbuareoon.t907 2)
B SynemamarcivDahl 1907) STE]
M Syemangtbelelesser 1919 |
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F Synemasimonesctessen 1919]
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
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274
Ansie S. Dippenaar-Schoeman et al.: South African spiders
' [ConSTATUS| eNOEMEITY PROV |EC|FS|6/KAN| LM) NC [NW | We
TgulbgenioasSmeon 85 —SSOS~STS
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Theriops meoopesDppenearScheenen 19893) SAE
Tromisospmetowss 1679 tt
Tronisopsseregaenas Mil 9@ St
Themiopssueaussimon 185 t t
TroniswausialsComeli 957 tT
Tromiasdorduskoscni080 AE
Troniascttnelssimen 875
Tremiascongoenis Comets 1957 tA
Troniowsdmsiteswen 919 tT
Troniasdaadolessimon 60
Troniowsgentouskasen 7000 Tt TT
Thomiseloloarrustawerce 19961) tt ta
Troniswsmechodoicomeini 98 Tt
Tromiasralestawence 922) THE
TromiasscbueelSmon 0) THE
Themis scopess(Smon8) tt
TromiasspeubsusFocock Ot)
TronisstonnglPocor 190A
Themis avaustawence 1928) RE NE
Tonsaticonstessent99 tt
Tmousconettomiswio.t92 ST tt ttt
TowscancelausTooeh 659)
Tmousconetinicaceneo,99 A
Tmovstoloustesen tse tA
Tmoweguneenoemion 9
Tonslongeoulausnilew 9
Towsratdersstesen 25 8 SAE
Tonsplontonssinen 3909
Trchopasismenitesimen 86
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Ptesovingtoweee 9 eae
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oles ratlersisLawence, 19982)
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Fetes tgdonsLawece 192 ——~SCS~*aY |
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 975
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘0 [Gow starus ENDaWCITY PROV [EC[FS/ 6 KN] LM] [Ww] wC
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B | Afroceto flabella Lyle & Haddad, 2010
M | Afroceto gracilis Lyle & Haddad, 2010
B_ Afroceto martini (Simon, 1897)
B | Afroceto plana Lyle & Haddad, 2010
M | Afroceto porrecta Lyle & Haddad, 2010
B | Afroceto rotunda Lyle & Haddad, 2010
B | Afroceto spicula Lyle & Haddad, 2010
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F | Capobula ukhahlamba Haddad, Jin, Platnick & Booysen, 2021
B | Fuchiba aquilonia Haddad & Lyle, 2008
B | Fuchiba capensis Haddad & Lyle, 2008 SAE
B | Fuchiba montana Haddad & Lyle, 2008 STHE
F Fuca sinilsHedéode yie2008 SSS) (OTSAE KING T
B FuchibatorsHaddod iyi, 2008 SOSA EOE
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B FuchbotuusbicomisHadéedsiye 2008 8) USE WET
F Fuchbotushaddediye,2013 6) OTSA SCE]
B FuchbotusKgelaHaddad eye 20082) STE
B JocquestuscapensstyeaHodded 2018 SSS «ODSSCSAESOWCES
F Jooqestusharisityle& Hedied2018 6 COOT SAE
B ooqestusincurs le Hades 2018 ia
B | Jocquestus roeweri (Lawrence, 1938) 5| pp | ‘sae | KZNE =
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
276
Ansie S. Dippenaar-Schoeman et al.: South African spiders
‘> [CoN STATUS] ENDEMIGTTY PROV EC|FS|G) KZN LM) NC | NW | We
M Mcheospustsuessen 1928] TT TT
F Miacheesscoplfersinonte56 8] OTS
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Playoldesabhotawence te 2) us| ome | | |)
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Playoldespusis Pocock 1058 AS
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F \Mptotesakemanwiti 964 | |S |] ttt) ||
F MogrammopesrevcaudusOP-Cambidge. 1682 2| te STE) TTT
F MogrammopesconsticusPucele08 | SAE TTT
B Miogrammopesiongcaudus OP-Cambidge, 10622) SHE
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F Philponelopercsa(Simont@56) 6 OTSA WE
F Uobous plnpedussimen 89 8| SAE
G Uobous pumpestucas 86 | TT
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Caesetius murinus Simon, 1893
Caesetius politus Simon, 1893 6 |
Caesetius schultzei Simon, 1910
Caesetius spenceri (Pocock, 1900)
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 277
Ansie S. Dippenaar-Schoeman et al.: South African spiders
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047
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279
Ansie S. Dippenaar-Schoeman et al.: South African spiders
Discussion and conclusion
South African spider systematics and ecology are in an exploratory phase and
the traditional approach to mapping diversity has enabled spider ecological re-
search results in South Africa to generate species lists that are often resolved
to species level. This descriptive phase provides the foundation for more inte-
grative work and any attempts to ignore the importance of providing baseline
diversity and taxonomic data will hamper subsequent attempts to develop a
deeper understanding and appreciation of this unique heritage. This checklist
provides a framework and context for future work.
Acknowledgements
We thank the ARC and SANBI Threatened Species Programme for technical
support; the Universities of Cape Town, Free State, Johannesburg, KwaZulu-Na-
tal, Limpopo, North West, Pretoria, Rhodes, Stellenbosch, Venda and Walter Si-
sulu and the Tshwane University of Technology for providing material; and the
National Research Foundation (NRF) for generous funding and support. The
staff of the Arachnology section at the NCA are thanked for their assistance
with sorting and databasing specimens collected during the SANSA surveys.
Various students, members of the public and the Spider Club of Southern Africa
collected material for SANSA. We also thank the South African National Parks,
E. Oppenheimer & Son and provincial conservation agencies for supporting and
providing the opportunity to collect in their parks and reserves and the provin-
cial conservation agencies for providing collecting permits.
Additional information
Conflict of interest
The authors have declared that no competing interests exist.
Ethical statement
No ethical statement was reported.
Funding
The National Research Foundation (NRF) Grant number 87311.
Author contributions
ASD conceptualized, compiled data and wrote the first draft; CRH data curation and
commented on drafts of the manuscript; RB data curation and commented on drafts of
the manuscript; LL data curation and commented on drafts of the manuscript; RS data
curation and commented on drafts of the manuscript; SHF data analysis and comment-
ed on drafts of the manuscript.
Author ORCIDs
Ansie S. Dippenaar-Schoeman ® https://orcid.org/0000-0003-1 532-1379
Charles R. Haddad © https://orcid.org/0000-0002-2317-7760
Ruan Booysen ® https://orcid.org/0000-0002-0841-9143
Stefan H. Foord © https://orcid.org/0000-0002-91 95-2562
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 280
Ansie S. Dippenaar-Schoeman et al.: South African spiders
Data availability
All of the data that support the findings of this study are available in the main text or
Supplementary Information.
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African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 288
Ansie S. Dippenaar-Schoeman et al.: South African spiders
Supplementary material 1
Reference list and citations for all the studies mentioned in the paper that
are in grey literature
Authors: Ansie S. Dippenaar-Schoeman, Charles R. Haddad, Leon N. Lotz, Ruan Booysen,
Rudolph C. Steenkamp, Stefan H. Foord
Data type: docx
Copyright notice: This dataset is made available under the Open Database License
(http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License
(ODbL) is a license agreement intended to allow users to freely share, modify, and
use this Dataset while maintaining this same freedom for others, provided that the
original source and author(s) are credited.
Link: https://doi.org/10.3897/Afrinvertebr.64.111047.suppl1
Supplementary material 2
List of Photo Identification Guides available from the World Spider
Catalogue or Zenodo
Authors: Ansie S. Dippenaar-Schoeman, Charles R. Haddad, Leon N. Lotz, Ruan Booysen,
Rudolph C. Steenkamp, Stefan H. Foord
Data type: docx
Explanation note: Each guide contains information on the genera and species recorded
from South Africa with information on global distribution, South African distribution with
a map, lifestyle, conservation and taxonomic information and images and photographs.
Copyright notice: This dataset is made available under the Open Database License
(http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License
(ODbL) is a license agreement intended to allow users to freely share, modify, and
use this Dataset while maintaining this same freedom for others, provided that the
original source and author(s) are credited.
Link: https://doi.org/10.3897/Afrinvertebr.64.111047.suppl2
African Invertebrates 64(3): 221-289 (2023), DOI: 10.3897/Afrinvertebr.64.111047 289
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